NOAA Technical Memorandum NMFS-F/NEC-19

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Northeast Monitoring Program

Environmental Benchmark Studies
inCascoBay — Portland Harbor, Maine,

April 1980

L!

Woods ]
Insti

U.S. DEPARTMENT OF COMMERCE

National Oceanic and Atmospheric Administration

National Marine Fisheries Service

Northeast Fisheries Center

Woods Hole, Massachusetts

January 1983

NOAA TECHNICAL (CMORANDUM NMFS-F/NEC

The Northeast Fisheries Center (NEFC) conducts targeted research to
provide needed information for fisheries resource and habitat managers in the
Northwest Atlantic. The NEFC operates facilities in Gloucester and Woods
Hole, Massachusetts; Narragansett, Rhode Island; Milford, Connecticut; Sandy
Hook, New Jersey; and Oxford, Maryland. It also administers the NMFS's
National Systematics Laboratory in Washington, D.C., and Atlantic
Environmental Group in Narragansett, Rhode Island. This targeted research
focuses on: (1) harvesting, aquaculture, and utilization of fisheries
resources; and (2) the health of the marine environment as it affects the
production and edibility of fisheries resources. Users of this information
include federal and state agencies, private industry, and the general public.

The real-time need for this information by decision makers often
precludes publication in formal journals. The NOAA Technical Memorandum NMFS-
F/NEC series identifies informal documents prepared by NEFC authors or similar
material prepared by others for NEFC purposes, where formal review and
editorial processing are not appropriate or feasible. However, documents
within this series reflect sound professional work and can be referenced in
formal journals. Any use of trade names within this series does not imply
endorsement.

Copies of this and other NOAA Technical Memorandums are available from
the National Technical Information Service, 5285 Port Royal Rd., Springfield,
VA 22161. Recent issues Of NOAA Technical Memorandum NMFS-F/NEC are noted
below:

4. Fishery Management Techniques, A Review. By Michael P. Sissenwine and
James E. Kirkley. October 1980. v + 10 p.

5. The Status of the Marine Fishery Resources of the Northeastern United
States. By Margaret M. McBride and Bradford E. Brown. December 1980. vii +
13 p., 4 figs., 3 tables.

6. Economic and Biological Data Needs for Fisheries Management, With
Particular Reference to the New England and Mid-Atlantic Areas. By Guy D.
Marchesseault, Joseph J. Mueller, and Ivar E. Strand, Jr. December 1980. v +
10 p. , 1 fig. , 3 tables.

7. Methodology for Identification and Analysis of Fishery Management
Options. By Brian J. Rothschild, Richard C. Hennemuth, Jacob J. Dykstra, Leo
C. Murphy, Jr., John C. Bryson, and James D. Ackert. December 1980. v + 10
p., 5 figs., 1 app.

8. Phytoplankton Community Structure in Northeastern Coastal Waters of the
United States. I. October 1978. By Harold G. Marshall and Myra S. Cohn.
August 1981. Revised and reprinted October 1981. v + 14 p., 4 figs., 1 app.

(continued on inside back cover)

NOAA Technical Memorandum NMFS-F/NEC- 19

This TM series is used for documentation and timely communication of
preliminary results, interim reports, or special purpose information; and
^ rctMosp ^r has not received complete formal review, editorial control, or detailed

_^^fl MJ^, editing

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Northeast Monitoring Program
NEMP III 81 B,D,G 0120

Environmental Benchmark Studies
in Casco Bay — Portland Harbor, Maine,

April 1980

Peter F. Larsen, Anne C. Johnson,
and Lee F. Doggett

Bigelow Laboratory for Ocean Sciences, West Boothbay Harbor, ME 04575

U.S. DEPARTMENT OF COMMERCE

Malcolm Baldndge, Secretary

* National Oceanic and Atmospheric Administration

xCZ^ * John V. Byrne, Administrator

c^^ cd National Marine Fisheries Service

"j ——_ ^f ■ r=K William G. Gordon, Assistant Administrator for Fisheries

Northeast Fisheries Center

Woods Hole, Massachusetts

January 1983

DOC

Woe.

In

TABLE OF CONTENTS

Foreword v

List of Figures vii

List of Tables xi

Introduction 1

Methods 6

Results and Discussion 10

Depth, Temperature and Salinity 10

Sediments 15

Sediment Carbon and Nitrogen 25

Trace Metals 31

Hydrocarbons 52

The Fauna 53

Density 53

Biomass 69

Species Per Station 78

Diversity 78

Classification Analysis 90

Summary and Tentative Conclusions 104

Acknowledgements 108

Literature Cited 109

Appendix 1 113

Appendix 2 11'

iii

FOREWORD

This issue of the NOAA Technical Memorandum NMFS-F/NEC series is a report
(Technical Report No. 22) prepared by the Bigelow Laboratory for Ocean
Sciences, under Contract No. NA-80-FA-C-00008 to the National Oceanic and
Atmospheric Administration's (NOAA) National Marine Fisheries Service
(NMFS). The contract was awarded as a part of NOAA's Northeast Monitoring
Program (NEMP), and was monitored by Mr. Robert N. Reid of NMFS's Northeast
Fisheries Center. The report was submitted to NEMP in September 1982 as a
final report on a benchmark survey of the benthos in the Casco Bay, Maine,
area.

The report has been reprinted virtually as submitted, with only minimal
changes in format. References to specific trade names in this report do not
imply endorsement by NOAA/NMFS.

John B. Pearce, Manager
Northeast Monitoring Program

LIST OF FIGURES
Figure Page

1. The location of Casco Bay 2

2. Key to maps of upper and lower Casco Bay 3

3. Some of the major existing facilities representing
threats to environmental quality in Casco Bay 4

4. Locations of the 56 benthic stations sampled in

Casco Bay, April 1980 7

5 . Station depths in feet 13

6. The annual temperature and salinity cycle at

Boothbay Harbor (Garfield and Welch, 1978) 16

7. The distribution of mean grain size in phi units
throughout Casco Bay 20

8. Regression of mean grain size in phi units against

depth in feet 22

9. The distribution of sand, silt and clay in the

Casco Bay samples 24

10. The distribution of organic carbon in the surficial
sediments of Casco Bay 26

11. The relationship between organic carbon and

bottom depth 29

12. The relationship between organic carbon and mean

grain size (phi units) 30

13. The distribution of cadmium (ppm dry weight) in

the surficial sediments of Casco Bay, Maine 33

14. The distribution of chromium (ppm dry weight) in

the surficial sediments of Casco Bay, Maine 36

VII

Figure Page

15. The distribution of copper (ppm dry weight) in

the surficial sediments of Casco Bay, Maine 38

16. The distribution of lead (ppm dry weight) in

the surficial sediments of Casco Bay, Maine 40

17. The distribution of nickel (ppm dry weight) in

the surficial sediments of Casco Bay, Maine 42

18. The distribution of zinc (ppm dry weight) in

the surficial sediments of Casco Bay, Maine 45

2

19. Distribution of density (individuals/m ) of

macrobenthos in Casco Bay, Maine 63

20. The relationship between density and bottom depth 66

21. The relationship between density and mean

grain size 67

22. Distribution of biomass (grams wet weight per

2
0.1m ) of macrobenthos in Casco Bay, Maine 70

23. Relative proportion of biomass of major faunal
components in Casco Bay 72

24. The relationship between biomass and bottom depth 75

25. The relationship between biomass and mean

grain size 76

26. The relationship between biomass and organic

carbon content of the sediments 77

27. The distribution of number of species per station
throughout Casco Bay 79

28. The relationship between species per station and

bottom depth 81

VJ.11

Figure Page

29. The relationship between species per station and

mean grain size 82

30. The relationship between species per station and

sediment organic carbon 83

31. The distribution of macrobenthic H' diversity in

Casco Bay , Maine 84

32. The relationship of H' diversity and bottom

depth 87

33. The relationship of H' diversity and mean

grain size 88

34. The relationship of H' diversity and sediment

organic carbon 89

35. Dendrogram of hierarchical classification

of stations in Casco Bay, Maine. Lower numbers
refer to station numbers. Numbers on branches
refer to site-groups 92

36. Distribution of site-groups in Casco Bay 93

37. Dendrogram of hierarchical classification of

species data. Letters refer to species-groups 95

38. Constancy of species-groups at site-groups 101

39. Fidelity of species-groups to site-groups 102

IX

LIST OF TABLES
Table Page

1. Location, depth, bottom temperature and salinity

of stations sampled in Casco Bay, April 1980 11

2. Sediment characteristics, organic carbon and

Kjeldahl nitrogen values for Casco Bay stations 17

3. ANOVA table of multiple regression analysis of
organic carbon on mean grain size, bottom depth

and their interaction 31

4. Concentrations of metals (ppm dry weight) in surface
sediments of Casco Bay, Maine 32

5. Correlation matrix for Casco Bay trace metal samples.... 47

6. Comparison of trace metal levels at several

New England locations 49

7. Phylogenetic listing of taxa encountered during benthic

sampling of Casco Bay, April 1980 54

2 2

8. Species per station, density/m , biomass (g/m wet

weight), diversity (H 1 ), evenness (J') and species

richness at each of the 56 Casco Bay stations

sampled 60

9. Mean density of invertebrates in unconsolidated
sediments of temperate and boreal inshore waters
(modified from Maurer et al . 1978) 68

10. Membership of species-groups 97

11. Mean, ranges and standard deviations of various

physical and biological parameters by site-group 105

12. Patterns of significant differences between site-groups
based on Duncan's multiple range test 106

XI

INTRODUCTION

Casco Bay, a large, complex bay, located on the south central coast
of Maine, is noted for its scenic beauty as well as for its importance
to business and commerce (Fig. 1). Within the approximately 400 square
kilometers comprising Casco Bay are 300 kilometers of coastline and
upwards of 400 islands (U.S. FWS , 1980). Included within Casco Bay is
the city of Portland, the largest in Maine, which ranks as one of the
busiest ports in New England, largely due to heavy petroleum traffic.
Portland is also the largest fishing port in Maine. Presently, 27% of
the coastal population of Maine is situated on Casco Bay. The growth of
this segment of Maine's population will be accelerated by the increased
use of Portland Harbor. Current expansion projects include a major ship
building facility, a fish pier, and a containerized cargo dock. Some of
the major existing facilities representing potential threats to
environmental quality are located in Fig. 3. At the same time that
human and industrial density is increasing in the Casco Bay region,
seals, eagles, black guillemots, and other species indicative of a clean
or undisturbed environment, are still found and the area remains heavily
utilized for commercial fishing.

In spite of the potential for conflict between development and the
traditional use of the nearshore waters for commercial fishing and
recreation, little systematic environmental evaluation has been
accomplished in Casco Bay. Hulburt (1968, 1970) and Hulburt and Corwin
(1970) investigated several aspects of the physical oceanography and
phytoplankton of the region while Jones (1980), and Parker and Garfield
(1981a, b) provided background information on microplankton production

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Fig. 2. Key to maps of upper and lower Casco Bay.

Fig. 3. Some of the major existing facilities representing

threats to environmental quality in lower Casco Bay.

7010"

43°40

-4340

Map A

70° 10'

X Oil and gasoline tank farm
[A Dredged area

70°I0'

70°00'

43 c
50'

N

Freeport municipal sewage discha

Port Clyde Packing Co. industrial-
discharge and Yarmouth municipal
■■•;'.'-^rt- • sewage discharge &v u 33

Oil storage
facility

Power station cooling
:;' " water discharge

■Gr.Chebeague?
i Island

05

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010

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2

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3
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Nautical Miles

40'

70° 10'

Map B

70°00'

and hydrography of Casco Bay, respectively. To date, the most
comprehensive faunal records are from the early surveys of Verrill
(1874) and Kingsley (1901). Few quantitative studies of the
shallow-water marine benthic communities in the boreal zone of the
eastern United States exist (Dexter, 1944, 1947; Hanks, 1964; Shorey,
1973; Bilyard, 1974; Larsen, 1979), and none consider the Casco Bay
region.

In April 1980, we undertook a broad scale benthic survey of Casco
Bay for the purpose of establishing an environmental benchmark against
which subsequent natural and man-induced fluctuations could be measured.
Due to the complex topography, hydrography and anthropogenic inputs,
great care was taken to insure that all possible variations in the soft
bottom habitat were included. Based on the results of this survey, a
long-term monitoring program of selected stations was instituted, and
the results of this effort will be presented in forthcoming documents.
This present report summarizes the physical and biological data from the
1980 broadscale survey.

METHODS

The basic sampling design involves four transects along the long
axis of Casco Bay with additional stations placed in areas of interest,
such as the major sounds between the islands and near potential point
sources of pollution. Station density is highest near Portland where
steep environmental gradients might be expected due to freshwater inflow
and more concentrated development. Station locations are presented in
Fig. 4.

70°I0'

43°40'

Nautical Miles

Peaks
Island

43 4d

Map A

Fig. 4. Locations of the 56 benthic stations sampled in
Casco Bay, April, 1980.

70° 10'

70°I0'

70°00'

43'
50'

N

>

Island

oio

o

2

_i_

3

4

_j

Nautical Miles

70° 10'

Mao B

70°00'

2
A single 0.1 ra Smith-Mclntyre grab sample was taken at each

station. Subsamples for sediment grain size, organic carbon and

Kjeldahl nitrogen analyses were removed from each grab. At 32 stations

additional subsamples were removed for heavy metal and hydrocarbon

analyses. In each case, prescribed procedures for the preparation of

subsample containers and the subsampling process were followed, and the

subsamples were frozen for delivery to the appropriate analytical

laboratory. Sediment grain size, organic carbon and Kjeldahl nitrogen

analyses were done by GEOMET Technologies, Inc., Melville, New York, and

heavy metal and hydrocarbon analyses were done by the National Marine

Fisheries Service. The main-body of the sample was sieved on nested 0.5

and 1.0 mm screens. The debris remaining on the screens was fixed in 5%

buffered formalin and returned to the laboratory for faunal analysis.

Bottom temperature and salinity were determined at each station using a

Beckman RS5-3 portable salinometer.

In the laboratory all organisms were transferred to 70% ethanol,
removed from the 1.0 mm size fraction and identified to the lowest
taxonomic level possible. Wet weight biomass was determined for the
major taxonomic groupings.

All data were entered and processed by the University of Maine
Computer Center through the Bigelow Laboratory Computer Center. Data
analyses included informational diversity and its components, calculated
by standard formulas given by Margalef (1958) and Pielou (1970), and
numerical classification in both the normal and inverse modes. The
Canberra metric dissimilarity index and the flexible sorting clustering
strategy were used in the latter procedure because of their demonstrated
success in marine benthic studies. The data were log transformed.

10

Nodal analysis, for both constancy and fidelity, was applied to

interface the results of the two classifications. Following the

convention presented in Boesch (1977), a constancy index was calculated

as C . . = a . ./(n.n .). where a., is the number of occurrences of
^J id *- 3 W

Species-Group i in Site-Group J and n. and n. are the numbers of

entities in each group considered. The fidelity index was calculated as

F..= (a.. .n.)/(n. . a..), the symbols having the same meaning as
^J 1*3 3 3 3 3 t>3

above. Fidelity values less than 1 suggest a negative relationship and
values of over 1 a positive relationship between a species-group and a
site-group.

RESULTS AND DISCUSSION

Depth, Temperature and Salinity

The depths of the 56 stations sampled range from 7 to 140 feet,
i.e. 2-43 m (Table 1). All stations exceeding 70 feet in depth are
offshore of the outer islands and most inner Bay stations are in the
25-50 foot depth range (Fig. 5). Depths are measured by fathometer and
are not corrected for tidal stage.

Bottom water temperatures in April range from 2.9 to 6.3 C with
most stations being between 3.0 and 5.0 C. Statistical analysis of
water temperature and depth indicates that vernal warming of surface
waters is already well progressed by April. This is a very highly
significant relationship, p > .9999, with a correlation coefficient of
-0.60222. With the exception of two stations, salinity throughout the
Bay varied within the narrow range of 30.3 - 32.9 /oo (Table 1). The
two stations outside of this range are the relatively shallow stations
35 and 41 with salinities of 25.6 and 19.8 /oo, respectively. Heavy

11

Table 1. Location, depth, bottom temperature and salinity of stations
sampled in Casco Bay, April, 1980.

EX 8001 Apr

il 198C

1

Station Number

Latitude

Longitude

Depth (m)

Temp °C

Sal (o/oo)

2

43°37'

.97

70°09'

.34

30.5

3.6

32.3

3

43°39'

.69

70°05'

.82

33.6

3.7

31.9

4

43°40'

.88

70°04'

.70

33.6

3.6

32.1

5

43°41'

.06

70°02'

.57

42.7

3.4

32.9

6

43°43'

.68

70°00'

.64

19.8

2.9

7

43°45'

.81

69°59'

.56

11.3

4.0

31.6

8

43°47'

.00

69°58'

.54

15.3

3.7

32.1

9

43°37'

.54

70°11'

.91

16.8

4.2

31.0

10

43°39'

.15

70°08'

.40

38.1

3.6

32.1

11

43°40'

.95

70°08'

.26

24.4

3.1

32.7

12

43°42'

.11

70°06'

.66

20.4

3.3

32.5

13

43°45'

.02

70°04'

.94

14.6

3.8

32.2

15

43°46'

.47

70°01'

.83

17.1

3.5

32.3

16

43°47'

.20

70°01'

.13

15.3

3.7

17

43°48'

.84

69°59'

.83

11.3

4.8

18

43°38'

.61

70°15'

.73

13.7

4.6

31.1

19

43°38'

.98

70°15'

.18

13.7

4.5

31.0

20

43°39'

.48

70°14'

.56

10.4

4.4

31.1

21

43°39'

.69

70°14'

.41

7.6

4.6

30.5

22

43°39'

.93

70°13'

.79

12.2

4.9

30.6

23

43°40'

.16

70°13'

.62

7.6

5.2

30.4

24

43°40'

.55

70°13'

.30

18.3

4.7

31.0

25

43°41'

.11

70°12'

.68

9.2

4.3

31.1

26

43°41'

.75

70°12'

.08

9.2

4.2

31.5

27

43°41

'.97

70° 11

'.53

13.1

4.6

31.6

28

43°41

'.36

70°11

'.04

14.6

4.4

31.7

29

43°42

'.88

70°10

'.51

13.7

4.5

31.7

30

43°43

'.29

70°09

'.96

12.2

4.4

31.7

31

43°43

'.99

70°08

'.95

12.2

6.3

32

43°44

'.97

70°08

'.17

7.6

4.6

31.3

33

43°46

'.12

70°05

'.65

11.6

4.0

32.0

34

43°47

'.69

70°03

'.54

10.7

4.0

32.2

35

43°49

* .20

70°01

'.86

7.6

4.9

25.6

36

43°40

'.66

70°14

'.44

7.9

4.5

30.7

12

EX 8001 April 1980

Station Number Latitude Longitude Depth (m) Temp C Sal (o/oo)

37

43°41'.12

70°14'.51

2.1

4.9

30.3

38

43°41'.75

70°12\08

8.2

4.4

31.4

39

43°43\34

70°11'.94

10.7

4.3

31.5

40

43°44*.94

70°10'.00

13.1

4.4

31.6

41

43°47'.04

70°07*.15

7.3

5.7

19.8

42

43°48'.22

70°05'.98

6.1

5.2

31.6

43

43°50'.06

70°01'.99

7.6

5.9

30.3

44

43°40'.51

70°13'.84

5.5

4.5

31.0

45

43°40'.80

70°13*.30

5.5

4.4

31.1

46

43°39'.50

70°13'.95

7.6

4.4

31.2

47

43°39'.36

70°13'.09

16.8

4.5

31.1

48

43°38'.70

70°13'.35

9.2

4.5

31.0

49

43°38*.51

70°12'.86

15.3

4.4

31.2

50

43°37'.87

70°12*.41

21.4

4.6

30.6

51

43°39'.61

70°12*.01

16.2

4.3

31.2

52

43°39'.45

70°12'.75

15.3

4.4

31.2

53

43°39'.31

70°12'.19

15.3

4.4

31.2

54

43°39'.25

70°09'.26

32.0

3.6

32.3

55

43°41'.18

70°10'.97

25.6

4.0

31.7

56

43°41'.90

70°09'.83

13.7

4.4

31.4

57

43°41'.99

70°09'.60

14.3

4.3

31.5

58

43°49'.93

70°00'.46

2.1

7.8

20.1

BOTTOM DEPTH

13

431144 •

143*113' ..

M3«4E' ..

43»H1 ' ..

43*140' .:

43*39'..

W3*38

43*37*

70M6* 70M5' 70°m* 70°13' 70*12' 70M1' 70*10' 70* 9"

Fig. 5. Station depths in feet,

14

BOTTOM DEPTH

143«51'

M3«U9' .-

143«39'

70MO' 70« 8' 70' 6' 70° 1' 70« 2* 70« ■ 69' 58'

15

rains and the spring freshet of the Royal River presumably combined to
temporarily depress the salinity at these stations.

No long-term temperature and salinity records are available for
Casco Bay. Long-term records taken since 1906 at Boothbay Harbor, 30
kilometers to the east, however, are closely representative of Casco Bay
(N. Garfield, personal communication). The annual temperature-salinity
cycle based on monthly means of Boothbay Harbor is presented in Fig. 6.
During the period of 1950 to the present, surface water temperature at
Boothbay Harbor varied between the extremes of -2.3 C and 23.0 C, while
salinity ranged from 25.0 to 33.6 °/oo (W.R. Welch, personal
communication) .
Sediments

The sediments of Casco Bay are predominantly fine (Table 2) .
Graphic mean grain size (Folk, 1974) ranges from -0.305 to 8.471 on the
phi scale although the grand mean is in the fine silt range at 6.345.
Only 8 stations have mean grain sizes in the sand range while 34 are in
the silt range and 13 can be classified as clay. The sand stations are
in areas of tidally scoured bottoms, such as the main approach to Casco
Bay, or in areas recently dredged. Generally, coarser sediments are
found offshore and in outer Portland Harbor, whereas fine sediments are
characteristic of the central and upper part of the Bay (Fig. 7).

Regression of mean grain size in phi units against bottom depth in
feet demonstrates that a significant relationship exists between the two
(Fig. 8). The correlation coefficient of -0.3317 is significant at the
98% level. Remembering, in this and subsequent regressions, that since
the phi scale is an inverse measure, this regression indicates that
coarser sediments may be expected in the deeper portions of the sampling

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Ma V f I November

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April ^ /

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March y

February

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Salinity 0/00

Fig. 6. The annual temperature and salinity cycle at Boothbay Harbor

(Garfield and Welch 1978).

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20

143a iju'

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43 g ME' ..

43 8 U1 ' ..

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43' 39'..

43" 38'

143*37'

70 , 16 1 70M5" 70°m' 70°13' 70ME' 70M1' 70M0' 70' 9'

Fig. 7, The distribution of mean grain size in phi units
throughout Casco Bay.

21

GRPJN SIZE

U3«51 *

t43«U9' ..

143«U7*..

ua'us 1 ..

U3«U3'__

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EXB001 CflSCO BflT

22

LU

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25 50 75
BOTTOM DEPTH

100

150

Fig. 8,

Regression of mean grain size in phi units against
depth in feet.

23

area. In many coastal areas sediments become coarser with increasing
distance from shore and/or from river mouths because estuaries serve as
traps for fine sediments. Since depth, too, commonly increases with
distance from shore, a positive relationship between depth and coarse
sediments might usually be expected in coastal waters. Due to the
complex topography of the Casco Bay region, we cannot separate the
effects of depth, distance from shore, or even distance from river
mouths in regard to gross sediment parameters. We feel confident,
however, that all three factors are operative in controlling the grain
size distribution, hence the scatter around the regression line.
Because we cannot meaningfully quantify distance from shore or distance
from source, we use depth alone to represent a complex of related
factors.

Fifty of the 55 stations have very poorly sorted sediments
(Table 2). With the exception of station 9, a coarse sand station which
is moderately well-sorted, the remaining stations are either poorly or
extremely poorly sorted.

The percentage of sand, silt and clay were calculated for each
sample and are plotted in Fig. 9 (see also Appendix 1). This
presentation suggests that most samples contain approximately equal
proportions of silt- and clay-sized particles and the mean grain size is
largely a function of the amount of sand in the sample.

Consideration of grain size distributions and sorting coefficients
does not in itself fully characterize the finer sediments of Casco Bay.
Our experience in taking grab samples shows that the sediments are also
extremely soft, i.e. have a very low bearing strength. Repeated
lowerings of our specially modified grab often failed to obtain a

24

0)

.c

-p

c

•H

>1

<C

l-l

o

TJ

C

m

-p

r-l

•H

(A

„

'O

c

a

to

IM

o

•

n

c

o

H

-H

a

4J

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(0

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en

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>i

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fl

W

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•H

T3

<u

en

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en

in

25

sample, although it had obviously been fully immersed in mud. In these
situations, we were forced to move the station slightly to find a more
cohesive bottom. Even here, however, the sediments visually resembled a
loose gel. In these cases, the bucket screens offered little impediment
to the grab's penetration into the bottom, and many organisms were
caught on the screens. In essence, the sediments were so low in bearing
strength that the grab sank to an unknown depth with the surface layers
being sieved by the bucket screens in the process.

Timson (personal communication) observed a similar sedimentary
phenomenon in Muscongus Bay, Maine, while working with sidescan sonar.
He describes it as a nepheloid or gel layer which is intermediate
between the water column and bottom. We call it fluid mud and believe
it is caused by tidal currents which are strong enough to prevent
complete deposition of silt and clay-sized particles, but which do not
have a sufficient excursion to disperse them from the system. If this
is true, we would expect variations in the depth and extent of the layer
over a spring-neap tidal cycle. In any event, the phenomenon must have
a profound effect on the nature of the benthic community and is worthy
of additional study.
Sediment Carbon and Nitrogen

Sediment organic carbon values as determined by chromic acid
digestion are presented in Table 2 and Fig. 10. Values range from 1.5
to 44.5 mg/g dry weight with an overall mean of 25.2 mg/g. Station 36,
which exhibited 233.0 mg/g organic carbon due to a high proportion of
wood chips, is an extreme outlier and is excluded from organic carbon
numerical analyses. The station is located at the mouth of the
Presumpscot River on a deposit of wood chips resulting from past

26

ORGRNJC CARBON

43 mn»

143«U3*

43«42' ..

U3 § m • ..

43* HO' __'

43* 39*..

43"38'

43*37'

70M6" 70M5' 70°1V 70°13 r 70»12' 70M1' 70" 10' 70* 8*

Fig. 10. The distribution of organic carbon (mg/g dry sedi-

men

t) in the surficial sediments of Casco Bay.

27

ORGANIC CARBON

143«51'

143«U9* ..

143«U7'__

U3*(45 • __

SOI
U3«U3'

H3«U1»__

143*39

70«10* 70" 8' 70" 8' 70° «t' 70« 2* 70« 0* 68" 58'

28

disposal of paper mill wastes into the river. No other station is so
influenced.

Examination of Fig. 10 shows that organic carbon levels lower than
the average are generally confined to the approaches to Portland Harbor
and the offshore stations. Stations with organic carbon levels
considerably higher than average occur in a group in lower Casco Bay off
Portland and Falmouth and at scattered sites close to land in the upper
Bay. In an attempt to elucidate potential controlling factors of
organic carbon distribution, organic carbon levels are regressed against
bottom depth (complex factor), mean grain size and distance from
Portland, the principal population and industrial center. Significant
relationships exist for both bottom depth and mean grain size (Figs. 11
and 12). These relationships suggest that organic carbon levels
decrease with increasing depth and increase with decreasing mean grain
size. The correlation coefficients are -0.4756 and 0.7742,
respectively, and both are significant at the 99.9% level. We show
above, however, that bottom depth and mean grain size are themselves
related and therefore, it is not clear from this analysis whether
organic carbon level is influenced by one or both of these factors. By
employing multiple regression analysis, it is shown that organic carbon
level is significantly related only to mean grain size (Table 3) .

EXB001 CflSCO BflT

29

50

M5 --

yo

35

30

25

G

CX3

20

CJ

o

»— «

15

2

CL

CD

CD

10

5 -■

25 50 75
BOTTOM DEPTH ( FT >

150

Fig. 11. The relationship between organic carbon and bottom
depth.

EXB001 CflSCO BflT

30

50

U5 --

yo --

35 -

30 --

K~l

25 --

GC
CC
CJ

20 --

CJ
>— i

z
cr

CD
CO
CD

15 --

10 --

1 2 3
GRAIN SIZE

Fig. 12. The relationship between organic carbon and mean
grain size (phi units) .

Table 3. ANOVA table of multiple regression analysis of organic carbon
on mean grain size, bottom depth and their interaction.

31

Source

Degrees of Type IV F

Freedom Sum of Squares Value

PR>F

Mean Grain Size 1

Depth 1

Grain Size-Depth 1

Error 50

451.0196

8.5920

2.9443

2079.9867

10.84 0.0018
0.21 0.6515
0.07 0.7917

Total Kjeldahl nitrogen values are also presented in Table 2.
Values range from 0.038 to 1.294 mg/g. To date no attempt has been made
to interpret these results. They are included for completeness.

Trace Metals

Subsamples from 32 stations were analyzed by atomic absorption
spectrometry for the metals cadmium, chromium, copper, lead, nickel and
zinc. Results, as ppm dry weight, are presented in Table 4.

Cadmium is present in the sediments of Casco Bay in concentrations
ranging from 0.20 to 0.90 ppm with a mean value of 0.50 ppm. Highest
cadmium values occur in the Portland vicinity and at station 53, the
former domestic dumpsite for Peaks Island residents (Fig. 13). Lowest
cadmium levels are found at the offshore stations while the remaining
stations deviate little from the mean.

Chromium levels average 34.5 ppm and range from 5.85 to 55.0 ppm
(Table 4). The approaches to Portland Harbor exhibit the lowest

32

Table 4. Concentration of metals (ppm dry weight) in surface sediments
of Casco Bay, Maine.

Sta. No.

Cd

Cr

Cu

Ni

Pb

Zn

2

< 0.25

27.0

9.45

11.0

13.5

39.0

4

0.40

26.0

8.38

18.5

18.5

49.4

8

0.30

23.0

8.70

13.0

12.0

43.0

9

0.20

8.50

2.40

4.5

10.5

20.85

10

0.35

39.1

14.0

22.8

29.7

70.8

11

0.25

31.0

11.4

18.5

24.0

59.5

13

0.50

36.5

11.8

19.5

21.5

65.5

15

0.55

38.0

20.0

20.0

33.5

73.5

16

0.55

54.0

16.4

27.5

25.0

30.5

17

0.60

47.5

16.6

32.0

19.5

84.5

19

0.87

49.2

44.5

23.65

61.4

81.9

20

0.80

46.5

32.0

18.5

51.0

100.

21

0.59

36.6

25.5

22.87

45.0

90.1

26

0.60

55.0

19.7

22.5

35.0

89.0

29

0.50

50.0

16.3

20.0

29.5

74.5

32

0.65

40.0

15.8

22.0

21.5

66.0

34

0.50

49.4

15.8

23.7

20.2

71.67

36

0.90

10.8

13.8

6.60

59.0

80.0

37

0.75

34.5

19.2

14.0

35.5

83.5

41

0.40

31.0

13.1

21.0

16.5

61.0

42

0.55

43.0

14.8

23.0

20.5

68.0

43

0.55

50.4

16.1

24.4

19.0

73.8

46

0.45

26.0

15.0

14.0

30.5

70.5

47

< 0.25

21.5

9.90

12.0

9.0

36.0

48

0.30

18.0

10.2

9.35

22.5

44.5

50

0.45

5.8

4.45

5.75

16.5

21.0

52

0.60

34.5

20.2

20.5

35.5

80.5

53

0.80

44.0

22.6

9.05

—

87.0

54

< 0.25

23.5

7.95

12.5

18.0

41.0

55

0.30

20.5

8.70

11.0

17.5

40.5

56

0.55

43.0

17.0

23.0

32.0

81.0

57

0.45

41.5

14.6

16.0

28.0

64.0

CADMIUM

33

H3»i|ir

43*43* . r

43*4?* ..

4 3 • U 1 ■ ..

43*40* _;

43*39* ..

LI3*38

43*37*

■

v

\

Y

/

tf

^

0.50

/

}

\ y

L

-<

C.60

.7V"

o.so

O.60

5^0. a? ^X^<o. ** cf

0.30

+

0.80

">

V

0.45

o.eo

<:

0.45

0.55

x

y

i

<0.-2-5

< . 2-5 -

+

+

+

70*16* 70*15* 70°14* 70°13* 70*12* 70*11' 70*10' 70* 9*

Fig. 13. The distribution of cadmium (ppm dry weight) in
the surficial sediments of Casco Bay, Maine.

34

CADMIUM

H3«51 '

143«U9 % ..

t43«U7*__

43«U5* ..

U3 , U3 , __

H3««U • ..

143«39'

0.40

<U.2S

0.35

P

\ h

70MO* 70" 8' 70* 6' 70" 1' 70' 2' 70" ' BS^Q

• 5R*

35

chromium levels and the Portland area in general is characterized by low
to moderate chromium levels (Fig. 14). Stations in Portland Harbor
proper and in mid and upper Casco Bay generally have higher than average
values.

Sediment levels of copper in Casco Bay range from 2.40 to 44.5 ppm
with an overall mean of 15.5 (Table 4). Only five stations exhibit
copper levels of over 20 ppm. These are the three stations in Portland
Harbor and stations 52 and 53 (Fig. 15). The gradient of decreasing
copper levels down the Fore River is suggestive of an upstream source.
Once again, lowest copper levels are found in the sandy main shipping
channel into the Bay and at the offshore stations. Upper Bay stations
generally are close to the mean in copper concentration.

Lead concentrations in the sediments range from 9.0 to 61.4 ppm
with an average of 26.8 (Table 4). Its distribution is similar to that
of copper, i.e. high concentrations in Portland Harbor with a decreasing
gradient down the Fore River, low concentrations in the tidal channel
and at most offshore sites, and low to moderate values throughout the
remainder of the Bay (Fig. 16).

Unlike the other metals, nickel does not show a strong pattern in
its distribution (Fig. 17). Concentrations range from 4.53 to 32.0 ppm
with a mean of 17.6 ppm. The extreme high value occurs at station 17 in
upper Casco Bay. Low values are found principally in channel areas and
just offshore, but even this pattern is not as clearly developed as for
the other metals.

Zinc concentrations in Casco Bay sediments average 65.4 ppm and
range from 20.5 to 100.5 ppm (Table 4). Highest values occur in

CHROMIUM

36

M3»tm'

^•l^' ..

43'42 % ..

U3»m ■ ..

43*38

431

43*110* .:

43'39' ..

70M6* 70M5' 70 o m* 70°13' 70M2' 70M1* 70M0* 70 f 9*

Fig. 14. The distribution of chromium (ppm dry weight) in
the surficial sediments of Casco Bay, Maine.

37

CHROMIUM

143*51 '

«43«H9« ..

I43«in*__

70* 10* 70" 8* 70 1 6' 70° *' 70 1 2' 70« ' 69' 58

■ SR«

COPPER

38

U3«HU«

U3«y3' ..

43*42* ..

43*41 ' ..

43*40' .:

43*39' ..

43 . 38 ,

43*37'

70M6* 70*15' 70°14' 70°13' 70M2' 70M1* 70*10' 70' 9'

Fig. 15. The distribution of copper (ppm dry weight) in
the surficial sediments of Casco Bay, Maine.

39

COPPER

U3«51 *

i43«U9' ..

t43«U7*._

^•MS* ..

U3«U3*

H3«U1 ' ..

143139.

70" 10* 70" 8' 70* B* 70' V 70" 2* 70« ' B^SB'

LERD

40

!43«HH«

43 f y3' .-

H3»42' ..

!43»U1 ■ ..

143* 40' .;

43'39' ..

43*38'

U3»37'

70' 16'

Fig.

70M5* 70°m* 70°13' 70ME' 70M1* 70M0' 70* 8"

16. The distribution of lead (ppm dry weight) in
the surficial sediments of Casco Bay, Maine.

41

LEAD

U3«51 •

143«U9' ..

143«U7'__

U3««5'__

li3«U3'._

143" 41 '__

US^'

70MO* 70" 8* 70 1 B* 70° V 70« 2' 70« % 69" 58'

42

NICKEL

43Hj 4 «

43»H3' ..

43«42' ..

43'Hl ' ..

43*4 0* .:

43'39' ..

43*38'

43 , 37"

70* 16*

Fiq

70«

17

70°l , r 70°13' 70M2' 70* 11 ' 70' 1CT

The distribution of nickel (ppm dry weight) in
the surficial sediments of Casco Bay, Maine.

43

NICKEL

H3«51 '

U3«U9« ..

U3 , U7'._

U3«U5* .-

U3'U3'._

i43««il * ..

U3«39'

70M0* 70* 8' 70" 6' 70° «t % 70« 2' 70« • 6S*5Q"

44

Portland Harbor although stations with values well above the mean are
found scattered throughout the Bay (Fig. 18). Low values are again
grouped in the outer shipping channel and offshore of the islands.

whereas upland drainage may be an important source of metals
deposited in coastal sediments, it does not seem to explain the elevated
levels in the Fore River (called the Stroudwater River in its non-tidal
portion). The Fore River has a small drainage area (28 sq. miles)
relative to the two other principal rivers entering Casco Bay, the
Presumpscot (590 sq. mi.) and the Royal (142 sq. mi.), which show little
or no elevation of metal levels near their mouths. In addition, above
Portland, the Fore River is largely surrounded by tidal marshes and
residential developments serviced by municipal sewers which discharge
elsewhere. It seems likely, therefore, that the elevated metal levels
in Portland Harbor sediments result from anthropogenic introductions
within the harbor and the industrialized lower Fore River estuary.
Additional sampling above Portland will be required to prove this
hypothesis .

Linear correlations were computed for the six metals as well as
percent organic carbon and mean grain size on the phi scale (Table 5).
This analysis shows that, as demonstrated elsewhere (i.e. de Groot et
at., 1976), metal levels in Casco Bay surficial sediments are highly
correlated with fine grained sediment and levels of organic carbon.
That these factors are significant is a reflection of the large surface
area of fine-grained sediments and the sorptive capacity of many organic
compounds. These relationships help to explain the areal distribution
of the metals as, for example, the uniformally low levels encountered in
the coarse sediments in the main entrance to Casco Bay.

ZINC

45

43144.

43*113' ..

43*12' ..

43*m ■ ..

Ms«no' .:

43*33' ..

43*38'

43*37'

70*16' 70*15" 70°m* 70 q 13' 70*12' 70*11' 70*10' 70* 9'

Fig. 18. The distribution of zinc (ppm dry weight) in the
surficial sediments of Casco Bay, Maine.

ZINC

46

43*51 *

143" 49 » ..

^•in* ..

43«U5' ..

43^3'..

U3«H1 ' ._

143139.

70MO" 70" 8" 70 1 6' 70" «t' 70« 2* 70« 0* SS^a'

47

The six metals were also highly correlated to each other in terms of
their distribution and concentration (Table 5). The only exceptions to
this generalization are the correlations of nickel with cadmium and lead
with chromium.

Comparison of trace metal levels in Casco Bay sediments with levels
found in other recent New England investigations will help to put these
results into perspective. Five studies utilizing comparable methodology
are available for comparison. Lyons et at. (in press) examined trace
metal levels in five northern New England estuaries. They concluded
from sediment profiles that three of them, Machias Bay, Cape Rosier and
the Seabrook River estuary show little increase in trace metal

Table 5. Correlation matrix for Casco Bay trace metal samples

X grain
Org. C Cd Cr Cu Pb Ni Zn size

Organic Carbon 1 .000

Cadmium 0.762 1.000

Chromium 0.822 0.461 1.000

Copper 0.578 0.705 0.635 1.000

Lead 0.476 0.775 0.287 0.802 1.000

Nickel 0.675 0.319 0.826 0.466 0.175 1.000

Zinc 0.853 0.786 0.766 0.775 0.724 0.625 1.000

Mean Grain Size 0.774 0.495 0.761 0.528 0.470 0.813 0.775 1.000

n = 32 except for lead and organic carbon where n = 31.
Significant at 99% confidence interval if r > 0.449 for n = 32 and
r > 0.456 for n = 31.

43

concentrations over the past century and are probably representative of
pre-industrial levels. Two other estuaries, the Saco and Kennebec,
exhibit recent anthropogenic enrichment due to industrial and/or sewage
inputs. Armstrong et al. (1976) determined trace metal values of the
sediments of the Great Bay estuary which has been historically subjected
to industrial discharges. Lyons and Gaudette (1979) investigated
concentrations in Jeffreys Basin, a fine-grained depositional area off
the coast of southern Maine and New Hampshire. They concluded that the
relatively high levels found there are the result of fine-grained
sediment export from estuaries. Two southern New England estuaries, the
unpolluted Mystic River estuary and the impacted Branford Harbor, were
contrasted by Lyons and Fitzgerald (1980). Finally, Greig et al. (1977)
analyzed a large number of sediment samples from Long Island Sound, a
large, highly "urbanized" estuary. For purposes of comparison we have
used only their results from the eastern half of the Sound, stations
72-143, to avoid the overbearing influence of inputs from the New York
City area.

Trace metal levels at the 11 New England sites are contrasted in
Table 6. It is important to remember that trace metal distributions in
Casco Bay are very heterogenous and the mean values are only a gross
representation of the conditions in a given subarea. Cadmium levels in
Casco Bay compare favorably with the three other sites having reported
values. The mean value is close to that of the unimpacted Mystic River
estuary and considerably lower than the values reported for Branford
Harbor and eastern Long Island Sound. Casco Bay sediments appear to be
moderately enriched in terms of chromium. The mean concentration is
nearly twice that of the pre-industrial levels of northern New

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England estuaries, but an order of magnitude lower than the Saco and
Great Bay estuaries, both of which are highly enriched with chromium due
to tannery operations (Armstrong et al. 3 1976; Mayer and Fink, 1980;
Lyons et al. s in press).

Copper levels in Casco Bay are also elevated relative to the
non-industrialized estuaries and are comparable to the other impacted
sites with the exceptions of the Kennebec River estuary, Maine and
Branford Harbor, Connecticut. Long Island Sound is the only other site
from which nickel data are available and the mean value is much lower
than that of Casco Bay.

The mean value of lead in Casco Bay sediments is higher than that
of the four non-industrialized sites and Long Island Sound, but
generally lower than the other industrialized estuaries. Mean zinc
concentration, on the other hand, is only exceeded by that reported for
Jeffreys Basin.

These results show that trace metals are not distributed
homogenously in the Casco Bay region. Whereas a strong correlation
exists between metal concentrations and both mean grain size and organic
carbon concentrations, there is also a strong geographic pattern not
completely explained by these relationships or the location of
freshwater inputs. In general, high trace metal levels are found in the
Portland area, which includes the lower Fore River estuary, low levels
are found in scour channels, relatively low concentrations are
encountered at the offshore sites and moderate levels occur in the very
fine sediments of central and upper Casco Bay. In addition, four metals
exhibit a gradient down the lower Fore River estuary suggesting an
upstream addition.

52

Comparisons with other New England sites indicate, with the
possible exception of cadmium, that trace metal concentrations in Casco
Bay are elevated well above presumed pre-industrial levels. Mean values
of each of the other metals examined are comparable to levels reported
from other industrialized New England areas.

Realizing that trace metal concentrations from stations in the
Portland area are generally much higher than the mean, and that the mean
is reduced by low concentrations elsewhere in the Bay, it is concluded
that the sediments of Portland Harbor and the lower Fore River estuary
are impacted by trace metals. Sediment profile studies are needed to
put the present levels into a historical context.

Hydrocarbons

The 32 hydrocarbon subsamples are in the process of analysis.
These results will be integrated into our overall analysis as soon as
they are available. Preliminary results indicate high levels of one or
more groups of hydrocarbons in Casco Bay sediments.

Visual examination of the samples indicated very high levels of
sediment hydrocarbons in the Portland area. Indeed, a couple of samples
were extremely difficult to pick because hydrocarbons formed a film on
the picking trays, forceps and organisms. Oil was also observed at one
of the sandy stations that is presumably well-flushed.

53

The Fauna

2
The 56 0.1m grab samples sieved to 1.0 mm yielded 264 putative

species (Table 7). Two hundred and thirteen of these were identified

to the species level. The molluscs, annelids and arthropods were the

best represented groups accounting for 16.6, 42.0 and 26.1% of the

species respectively. Species list for individual stations are

presented in Appendix 2.

Many of the species occurrences are interesting in terms of their

presence or abundance. For instance, one of the dominant polychaetes,

Aglaophamus neotenus, was described only very recently (Blake, 1980).

Among the isopods, both the Virginian Cassidinidia lunifrons and the

Arctic Munna fabvicii were found within the limited confines of Casco

Bay. Undescribed members of the amphipod genera Melita and Monoculodes

were encountered as were individuals of the genera Bathymedon, Hattmedon

and Gitanopsis . These latter, and perhaps several other records,

represent range extensions which will be treated in another

contribution.

Density

2
Numbers of individuals ranged from 120 to 36,380/m with a mean of

2
8,743/m (Table 8). The lowest value occurred at station 36 in the wood

chip deposits while the highest value was recorded at the nearby station

37 which was located in a mussel reef. Density distribution is

2
presented in Fig. 19. Values exceeding 10,000/m are generally found at

the offshore stations and at several stations in the lower Bay.

Three regions of low density stations are noticeable. These are

stations 47, 49 and 50 in the main channel into the Bay, stations 18, 19

liable 7. Phylogenetic listing of taxa encountered during benthii
sampling of Casco Bav, April 1980.

54

PHYLUM CNIDARIA

Class Hydrozoa

Campanularia sp.
Sertularia pumila
Hydroid A
Hydrozoa

Class Anthozoa
Anemone A
Anemone B
Cerianthus borealis

PHYLUM PLATYHELMINTKES

Notoplana atomata
Platyhelminth.es

PHYLUM RHYNCH0C0ELA

Cerebratulus laateus
Li-neus ruber
Nemertea A
Nemertea B
Nemertea C
Nemertea D
Nemertea E
Nemertea F
Nemertea G
Nemertea H
Nemertea I
Nemertea J
Nemertea K

PHYLUM BRY0Z0A

Caberea ellisi
Membraniporidae

PHYLUM MOLLUSCA

Class Gastropoda

Alvania arenaria
Alvania oarinata
Calliostoma ooaidentale
Cooculina sp.
Cyliahna alba
Cyliahna gouldi
Do to ooronata
Hydrobia sp.
Lacuna vinata
Littorina littorea
Littorina obtusata
Nassarius trivittatus
Oenopota bioarinata
Philine finmarehia
Skeneopsis planorbis

55

Class Scaphopoda

Dentalium entale

Class Vivalvia

Anomia aculeata
Avctia islandica
Astarte borealis
Astarte undata
Bivalvia
Cardita borealis
Cerastoderma pinnulatum
CKlamys islandica
Crenella decussata
Gemma gemma
Lyons-la hyalina
Maooma balthica
Modiolus modiolus
Mulinia lateralis
Mya arenaria
Mytilus edulis
Nuoula annulata
Nuaula delphinodonta
Nuoula tenuis
Pandora gouldiana
Periploma leanum
Periploma papyratium
Pitar morrhuana
Solemya borealis
Tellina agilis
Tracia oonradi
Thyasira flexuosa
Yoldia lima tula

PHYLUM ANNELIDA

Class Polychaeta

Aglaophamus oiroinata
Aglaophamus neotenus
Ampharete aoutifrons
Ampharete arotica
Apistobranchus tullbergi
Aricidea jeffreysii
Aricidea quadrilobata
Arioidea suecica
Archiannelida
Asabellides oculata
Autolytus sp.
Brada granosa
Brada villosa
Capitella aapitata
? Chaetopterus sp.
Cirratulidae
Clymenella torquata
Diplocirrus hirsutus
Dodeoaoeria sp.
Eteone flava
Eteone heteropoda

56

Eteone longa

Eucylymene collaris

Eusyllis blomstrandi

Exogone hebes

Exogone verugera

Gattyana cirrosa

Goniada maculata

Harmothoe extenuata

Harmothoe imbvioata

Hartmania moorei

Heteromastus filiformis

Laoniae cirrata

Lepidonotus squamatus

Lumbrineris acuta

Lumbrineris brevipes

Lumbrineris fragilis

Lumbrineris tenuis

Maldane sarsi

Maldanopsis elongata

Mediomastus ambiseta

Melinna oristata

Microphthalmos aberrans

Myriochele heeri

Nephtys bucera

Nepktys ciliata

Nephtys inoisa

Nephtys sp .

Nereidae

Nereis grayi

Nereis pelagiaa

Nereis sp.

Nereis virens

Nereis zonata

Ninoe nigripes

Notomastus latericus

Ophelina acuminata

Ophioglycera gigantea

Owenia fusiformis

Oweniidae

Paraonis gracilis

Paraonis lyra

Parapionosyllis longocirrata

Petaloproctus tenuis

Pherusa af finis

Pholoe minuta

Phyllodoce groenlandica

Phyllodoce maculata

Phyllodoce mucosa

Phyllodocidae

Polychaete B

Poly cirrus eximus

Poly cirrus medusa

Poly cirrus phosphoreus

Polydora ligni

Polydora quadrilobata

Polydora socialis

Polydora sp.

57

Potamilla neglecta
Praxillella gracilis
Praxillella praetermissa
Praxillella sp.
Prionospio steenstrupi
Pygospio elegans
Rhodine loveni
Sabella penicillus
Saalibregma in f latum
Sooloplos robustus
Scoloplos sp.
Sphaerodoropsis minuta
Sphaerosyllis erinaoeus
Spio filicornis
Spio setosa
Spiophanes bombyx
Spirorbis borealis
Spirorbis sp.
Stauronereis caecus
Stauronereis rudolphi
Sternaspis scutata
Streblospio benediati
Syllidae
Syllis cornuta
Syllis gracilis
Terebellid A
Terebillid B
Terebellidae
Terebellides stroemi
Tharyx sp.

Trichobranohus glaoialis
Trochochaeta multisetosa

Class Oligochaeta
Oligochaeta

PHYLUM SIPUNCULA

Golfingia verrillii
Phascolion strombi
Phascolopsis gouldii

PHYLUM ECHIURIDA

Eahiurus echiurus

PHYLUM ARTHROPODA

Subclass Cirripedia

Balanus balanoides

Subclass Malacostraca
Order Cumacea

Campylaspis rubicunda
Diastylis abbreviata
Diastylis cornuifer
Diastylis polita

58

Diastylis quadrispinosa
Diastylis sculpta
Eudorella hispida
Eudorella truncatula
Leptostylis longimana
Oxyurostylis smithi
Petalosarsia declivis

Order Isopoda

Cassidinidea lunifrons
Chirodotea ooeaa
Edotea triloba
Jaera sp.
Lirrmoria lignorum
Munna fabricii
Ptilanthura tenuis

Order Mysidacea

Erythrops erythrophthalma
Meterythrops robustus
Mysis stenolepis
Neomysis americana

Order Amphipoda

Aeginina longiaornis
Ampelisca abdita
Ampelisca agassizi
Ampelisca macroeephala
Ampelisca vadorum
Anonyx liljebovgi
Argissa hamatipes
Bathymedon sp.
Byblis gaimavdi
Capvella unica
Casco bigelowi
Corophium crassicorne
Corophium insidioswn
Corophium tuberculatum
Corophium volutator
Dexamine thea
Dulichia monacantna
Erichthonius rubricornis
Gammarus oceanicus
Gitanopsis sp.
Halimedon sp.
Haploops tubicola
Harpinia propinqua
Hippomedon serratus
Leptocheirus pinguis
Mayerella limicola
Melita n. sp.
Metopella angusta
Monoculodes n. sp.
Monoculodes tesselatus
Orchomenella pinguis
Paracaprella tenuis
Photis macrocoxa
Phoxocephalus holbolli
Pleustes panoplus
Pleusymtes glaber

59

Pontogeneia inermis
Protomedeia fasciata
Psammonyx nobilis
Stenopleustes gracilis
Stenopleustes inermis
Unioola irrorata
Order Decapoda
Cancer borealis
Cancer irroratus
Pagurus arcuatus
Pagurus longicarpus
Pagurus pubescens

PHYLUM ECHINODERMATA

Class Holothuroidea
Chirodota laevis
Molpadia oolictica

Class Echinoidea

Echinarachnius parma

Strongy locentrotus droebachiensis

Class Stelleroidea

Amphipholis squamata
Asterias sp.
Ophiopholis aculeata
Ophiura sarsi

PHYLUM HEMICHORDATA

Saccoglossus kowalevskii
Stereobalanus canadensis

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DENSJTY

63

U3»im •

U3 , 113'._ — ^ j

M3'42 % ..

U3«41 ' ..

143*40' .:

43'39' ._

Z\l

M3'38

43*37'

I see

356G.

loao.

^

19830.

2B330. --

70M6* 70M5' 70°m* 70°13' 70M2' 70" 11" 70M0' 70* 9*

Fig. 19. Distribution of density ( individuals /m ) of macro-
benthos in Casco Bay, Maine.

d e n s : T y

64

M3*51 '

43°U3 ■ ..

liiuo.

^

r

1

as

1

i 272

M3 tf U7 ' 1

x^r

V-V

■> /

■-9^2. .

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75!3

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8080. ^

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9250.

/7

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6080.

1080
U3«U3 * ._

205(1.

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9 I

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43'39'

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26470.

65 60. //>/

(7

13920

8180.

+

+

70M0* 70 1 8* 70» 6' 70° >4 ' 70* 2* 70« 0* SS'SS'

65

and 20 in Portland Harbor and several stations in the middle and upper
Bay. Reasons for these low densities are not completely obvious.
Stations with similar sediments located around stations 47, 49 and 50
have much higher densities. Further data are needed before we can
speculate on whether this is a natural or pollution-induced phenomenon.
Stations 18, 19 and 20 exhibited the highest levels of trace metals, so
perhaps the reduced densities there are impact related. The middle and
upper Bay stations with low animal densities are those where extremely
soft sediments were encountered. We believe that these sediments offer
so little bearing strength that only a depauperate community can
develop.

Correlation analysis was used to add insight into factors that
might be influencing density levels. Density is correlated with both
depth and mean grain size (Fig. 20 and 21). These relationships, both
of which are significant at the 99%, level indicate that density
increases with increasing depth and decreases with decreasing mean grain
size. The latter relationship adds support to the hypothesis that
physical properties at some fine-grained stations prevent the
development of a normal community. Density is not significantly
correlated with temperature, salinity or organic carbon content.

Correlation analyses were also run between density and the six
trace metals. Two of the metals, chromium and nickel, were negatively
correlated to density at over the 95% level. While not attaining the
conventionally accepted 95% level of significance, it is interesting to
note that cadmium, zinc, copper and lead are all negatively correlated
with density. The significance levels are 94, 93, 88 and 76%,
respectively.

66

EX8D01 CflSCO BAY

40000

35000 --

30000 --

25 50 75 100 125 150

BOTTOM DEPTH (ft)

Fig, 20. The relationship between faunal density and bottom
depth.

EX8001 CflSCO BAY

67

40000

35000 --

30000 --

GRAIN SIZE

Fig. 21. The relationship between faunal density and mean
grain size.

68

Comparing Casco Bay faunal densities with those from other
temperate and boreal areas demonstrates the comparative richness of the
region (Table 9 ). Such high density and biomass (see below) indicates
a high or extended period of productivity. Data on primary productivity

Table 9. Mean density of invertebrates in unconsolidated sediments of
temperate and boreal inshore waters (modified from Maurer et
at. 1978)

Location

Mean Density/:

Source

Casco Bay, Maine 8,743
Sheepscot Estuary, Maine

Gradient Study 4,928

Shallow Water Study 771

Mystic River, Massachusetts 3,000

Moriches Bay, New York 1,300

Delaware Bay 722

False Bay, South Africa 2,200

Gullmars Fjord, Sweden 4,198

Lambert Bay, South Africa 1,153

this study

Larsen & Doggett (1978)
Larsen (1979)
Rowe et at. (1972)
O'Connor (1972)
Maurer et al . (1978)
Field, (1971)
Rosenberg (1973)
Christie (1976)

of Casco Bay are presently being generated by other researchers at the
Bigelow Laboratory. These data may provide an explanation for the
observed faunal densities.

69

Biomass

2
Biomass of the 1.0 mm sieve fraction averaged 49.6 g/m on a wet

2
weight basis. The range at individual stations was 1.8 to 191.0 g/m

(Table 8) . In all cases animals weighing over one gram were excluded

from the analysis. Annelids constituted 49.6% of the fauna in terms of

wet weight. Arthropods, molluscs, echinoderms and miscellaneous phyla

accounted for 19.3, 11.6, 1.9 and 17.6% of the biomass, respectively.

There was considerable variation in total biomass between stations
but relative dominance of higher taxa was fairly consistent (Fig. 22 and
23). Annelids were biomass dominants at 43 of the 56 stations.
Arthropods were dominant at stations 2, 4 and 5 due to the abundance of
Ampelisoa agassizi and Haploops tubiaola and at stations 24 and 25 due
to Casoo bigelowi. Molluscs were also biomass dominants at only five
stations. These were stations 47 - 50 because of the presence of Nuoula
delphinodonta and Nassarius trivittatus and at the sparsely populated
station 43. Stations 18 - 20 in Portland Harbor were dominated in terms
of biomass by various miscellaneous taxa.

Like density, biomass exhibits a strong positive correlation
(99.9%) with depth and a strong negative correlation (99.0%) with mean
grain size (Figs. 24 and 25). Furthermore, it is negatively correlated
with organic carbon (99.9%) (Fig. 26). In addition, biomass is also
positively correlated at the 99.9% level with density and number of
species per station. It is negatively correlated with all of the trace
metals. Three of these relationships, cadmium, chromium and zinc are
significant at the 95% level. Levels of significance for copper, nickel
and lead are 83, 89 and 79%, respectively.

BI3MRSS

70

43 , H'4

U3' H3 f

us^e* ..

49*m • ..

43*40* .:

/■

43'39* ..

43' 38'

43 , 37'

19.1 --

70M6" 70M5" 70°14' 70"13' 70M2' 70M1* 70M0* 70 f 8'

Fig. 22. Distribution of biomass (grams wet weight per 0.1
m2) of macroben t hos in Casco Bay, Maine.

71

BIQMASS

U3«51 '

43«U9' ._

143«U7'__

U3«ys %

U3«U3*..

U3«U1 * ..

143«39*

70« 10' 70» 8' 70* 6' 70° «t* 70" 2' 70« * 69«58'

72

20

17.5 _

15 .

12.5 _

2 10 .

03

(D
J

I 7.5

o

k_

5 .

2.5 _

Miscellaneous

Molluscs

Crustaceans

y& Polychaetes

2 3 4 5 6 7 8 9 10 II 12 13 14 15 16 17 18 19 20

STATION NUMBER

Fig. 23. Relative proportion of biomass of major faunal components
in Casco Bay .

73

17.5 .

S

| Miscellaneous
Mol luscs

Crustaceans
Polychaetes

<

ffl raffl

n

21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39

STATION NUMBER

74

40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58

STATION NUMBER

EX8001 CflSCO BAY

75

en
<n
cr

in'
i— «

25 50 75

BOTTOM DEPTH (ft)

100 125

150

Fig. 24. The relationship between biomass and bottom depth.

EX8001 CflSCO BflT

76

en
<n
a.

ED
•—i
CD

123U56789
GRR1N SIZE I

Fig. 25. The relationship between biomass and mean grain size.

77

EXB001 CflSCO BflT

en

1— 1

10 15 20 25
ORGRNIC CflRBON

Fig. 26. The relationship between biomass and organic carbon
content of the sediments.

Species Per Station

2
The number of species per station (0.1m ) ranged from 5 to 86 with

a mean of 33.1 (Table 8). The offshore stations consistently had the

highest numbers of species and moderate numbers are characteristic of

lower Casco Bay. Less than 20 species were found at many of the

finer -grained stations and in the wood chips of station 36. Indeed,

stations 17, 32, 35, 36 and 43 were occupied by less than 10 species

(Fig. 27).

The number of species per station is very strongly correlated
(99.9% level) with increasing depth (Fig. 28). It is also negatively
correlated at the 99.9% level with mean grain size and organic carbon
(Figs. 29 and 30). Number of species is the only biological parameter
to be significantly correlated with salinity (98% level) .

As with density and biomass, species per station is negatively
correlated with each of the metals. Two of these relationships, cadmium
and zinc, are significant at the 99% and 95% levels, respectively.
Other non-significant correlations and their significance levels are:
chromium 94%; nickel 93%; copper 82% and lead 75%.

Diversity

Informational diversity, as measured by the Shannon index, ranged
from 0.415 at station 32 to 4.347 at station 10 (Table 8). The overall
mean was 2.72. High values of H' diversity are found at some offshore
stations and generally throughout the Portland region including the
trace metal impacted, low density stations 18-20 (Fig. 31). An
explanation for this unusual result can be found in an examination of
evenness and species richness levels (Table 8). In general, the high

NUMBER OF SPECIES

79

143144'

43 , 39* ..

43*38'

U3»37'

59. --

70M6" 70M5' 70°m' 70°13' 70M2* 70M1' 70MCT 70« 8"

Fig. 27. The distribution of number of species per station
throughout Casco Bay.

80

NUMBER OF SPECIES

U3«51 '

H3«U9* .-

U3««il ' ..

43139.

70* 10' 70« 8* 70' 6' 70° V 70« 2* 70* ' OS'SB*

81

EX8001

CflSCO BAY

LU

•— i

CJ
LU

cn

or.

LU
CD

s:

00 -
90 -

1 1 1

— 1

X

— 1 —

--

80 -

yS ~~

70 -

X

/*

60 -

X

X /

X

m m

50 -

X

x -X

40 -

--

30 -

20 -

10 -

-

I

M X/^ K

X / X *

IK «* ■»

/ • ^"* X

v«

>* X

X

1 1 1

— 1

1

--

25 50 75

BOTTOM DEPTH (ft)

100

125

150

Fig

28. The relationship between species per station and
bottom depth .

EXB001 CRSCQ BRT

82

100

on

UJ

•— i

CJ

UJ

a_

en

CL
UJ
CD

1 2 3
GRAIN SIZE

7 8

Fig. 29. the relationship between species per station and
mean grain size.

EXB001 CflSCO BRT

83

100

UJ

•— •

CJ
LxJ
Q_

tn
u.

CD

CL
Ul

m

Fig

10 15 20 25
ORGRNIC CARBON

30. The relationship between species per station and
sediment organic carbon.

DIVERSITY

84

43«UU' 4-

43*143' ..

43*42*. _

43*41 ' ..

43*40' .:

43*39' ..

43*38'

43*37'

70*16' 70*15' 70°14' 70°13' 70*12* 70*11' 70*10* 70* 3'

Fig. 31. The distribution of mac roben t hie H' diversity in
Casco Bay, Maine.

DIVERSITY

U3«51 *

U3«U9* ..

t43«in*._

U3«US' ..

U3«U3 % ._

H3«41 ' ..

43«39'

70MO* 70" 8' 7Q» 6' 70° «t* 70« 2' 70« 0' B^SO'

86

diversity values at many stations seems to be caused by high species
richness. Naturally, these are the stations with high numbers of
species. At other stations, however, evenness appears to be the
dominant component of diversity. For example, stations 15, 16, 18, 19,
20 and 36 only have moderate to low species richness levels but are
among the highest stations in evenness which results in diversity levels
of over 3.0 at stations where conventional logic would predict depressed
diversity.

Extremely low diversity is limited to stations with very fine
sediments. Not all the fine-grained stations exhibit such low
diversity, however, and the explanation for this is analogous to that
presented above.

Like the other biological parameters discussed, informational
diversity is positively correlated with bottom depth, at the 95% level,
and negatively correlated with mean grain size and organic carbon, both
at the 99% level (Figs. 32, 33 and 34). As would be expected, diversity
is positively correlated at the 99.9% level with species number, but
shows no relationship to density. Additionally, diversity is not
significantly correlated with any of the trace metals and is not even
consistent in the sign of the correlation coefficient.

Whereas, we believe all of the biological parameters should be
re-evaluated in greater depth once the hydrocarbon data can be factored
into the analysis, all of them, except diversity, presently add insight
into the existing conditions in Casco Bay and will aid us in providing
an integrated overview of the biological functioning and health of the
system. Diversity, on the other hand, is not presently useful in this
regard. Once all the data are available, diversity should again be

EXB001 CflSCO BAY

87

25 50 75

BOTTOM DEPTH (ft)

100

150

Fie. 32. The relationship of H' diversity and bottom depth,

EXB001 CflSCQ BRT

88

5.0

H h

H \-

5 L -°

0.5 --

0.0

<9

H 1 1 1 1 h

-1

1 2 3
GRRIN SIZE

6 7 8 9

Fig. 33. The relationship between H' diversity and mean
grain size.

EXBD01 CflSCO BAY

89

5.0

4.

5 --

4.

--

3.

5 -*

3.

--

2.

5 --

>-

h-

i— i

01
CO

Lu
>
i— i

o

2.

i

1

--
.5 --

,0 --

0.5 --

0.0

5 10 15 20 25 30 35 UO U5 50
ORGANIC CARBON

Fig. 34. The relationship of H' diversity and sediment organic
carbon .

90

evaluated, but other related tools, such as fit to a lognormal
distribution, should be looked at as well. It may be that H* diversity
is simply not the appropriate index to illuminate the finer nuances of a
heterogeneous system like Casco Bay.

Classification Analysis

Classification or cluster analysis is a useful way of objectively
examining patterns in complex data sets which cannot easily be uncovered
by other techniques. It is a hypothesis generating technique which
can suggest relationships between biological and physical factors that
may be causal to observed community distributions. Although the method
is numerical, and therefore objective, the interpretation is subjective.
One way to minimize subjectivity is to produce a large number of
site-groups and species groups and then combine them until the most
meaningful pattern is produced using a minimum of groups. The goal is
to produce the most comprehensive but simple explanation for the
observed phenomena.

We are at an intermediate step in this process. We have produced
station and species dendrograms based on faunal data. We have defined a
moderate number of site-groups and species-groups and have initiated a
comparison of the groups with the extrinsic factors now on hand. In
order to conserve resources we intend to wait until all of the physical
data are available before finalizing our interpretation. We have,
however, included our analysis to date because it does provide useful
information about Casco Bay and illustrates the strength of the analysis
that will be available soon.

91

The dendrogram which resulted from the classification of the 56
stations using species abundances as attributes (normal classification)
is presented in Fig. 35. For the time being we have truncated this
dendrogram at the nine group level. Examination of Fig. 35 shows that
all of these groups are fairly discrete but some, for example groups 2
and 3, are candidates for further fusion. We have a great deal of faith
in this classification because it shows good spatial discrimination
(Fig. 36).

The 10 members of site-group 1 are principally deep-water offshore
stations. Site-groups 2, 3 and 7 are limited to the Portland region.
Stations in site-groups 2 and 3 are intermingled in outer Portland
Harbor and are adjacent to one another on the dendrogram. This suggests
a close faunal affinity between them. Site-group 7 stations are found
on the edge of the patch of site-group 2 and 3 stations and are far
removed from them in the dendrogram. This reflects a real difference in
faunal composition undoubtedly controlled by physical factors.
Site-group 4 is widely scattered throughout Casco Bay with all but one
of the member stations occurring near shore. Site-group 5 dominates the
central portion of the Bay while site-group 6 members ring the Bay at
shallow stations. The three member site-group 8 exhibits no spatial
pattern and site-group 9 is a single station outlier consisting of the
mussel reef comminity at station 37.

The dendrogram resulting from the inverse classification is
presented in Fig. 37. Only those noncolonial species occurring at over
10% of the stations were used in this procedure. We have tentatively
truncated this dendrogram at the 14 group level. The most significant
feature of this analysis is the distinct separation of species-group N

92

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O

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o

o

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C\J

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<J-

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6

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<u

6

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c

c
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IH 4-1
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c o

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a.

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h ai oo
u

4-1 (-1

CO 01

<4-l

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CO CO 4J

CO (-1 -H

CO 0) CO

O g O

3 4-1

a

U _)

u cu

CU CM

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CD

(-1

CU

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CU
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c
co

1-4

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g - c

CO >, O
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O CU
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CO g

aj co 3

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— i —
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d

60
•H
fa

93

70°I0'

43 < 40'

S> Peaks
Island

Nautical Miles

4340'

70° 10'

Fig. 36. Distribution of site-groups in Casco Bay

94

70°I0'

70°00'

43'
50'

N

- .

&*j£ Peaks
Island

o

2

4

Nautical Miles

40*

70° 10'

70°00'

95

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96

from the others. This group is not a single specis outlier, but the
largest group and the separation suggests a basic difference in
distribution between species-group N members and members of the other
groups. The membership of each species-group is presented in Table 10.

By examining the constancy and fidelity of species-groups at the
various site-groups it is possible to achieve insight into the
distribution of the species-groups and perhaps into the controlling
ecological mechanisms. This process is called nodal analysis.

The patterns of constancy and fidelity of the species-groups at the
site-groups is summarized in Figs. 38 and 39. The width of the rows and
columns is proportional to the size of the groups. Site-group 1 is
occupied in medium to very high constancy by all species-groups with the
exception of species-groups H-K. These latter four groups also
demonstrate a fidelity of less than unity at site-group 1 indicating an
avoidance of the member stations. It is the only site-group where
species-groups E, F and M are highly constant and G and M are highly
faithful. With the exceptions of species-groups F and G, site-groups 2
and 3 are occupied by similar species-groups but differ in relative
constancy and fidelity especially in terms of species-groups A, B and C.
Site-group 4 is best characterized by the presence of species-groups I
and N. Species-group I is highly constant and faithful only at
site-groups 4 and 5. These two mid-Bay site-groups differ from one
another in that site-group 4 has six species-groups present at low to
medium constancy which do not occur at site-group 5 stations.

Site-group 6 is impoverished. Only species-group J is present at
moderate constancy. Site-group 7 has similarities to site-groups 2 and

97

Table 10. Membership of species-groups

Species -groups

Species-group A

Cerianthus borealis
Crenella deoussata
Periploma papyratium
Thyasira flexuosa
Eteone longa
Pherusa affinis
Pholoe minuta
Sabella penicillus
Phoxoeephalus holbolli

Species-group B

Modiolus modiolus
Mya arenaria
Nuaula annulata
Pi tar movvhuana
Ampharete aoutifrons
Stauronereis oaeous
Stenopleustes inermis

Species-group C

Cevastodevma pinnulatum
Paraonis gracilis
Harpinia propinqua
Orohomenella pinguis
Phoxis maoroaoxa

98

Species-group D Nemertea C

Phyllodoce mucosa
Casco bigelowi
Leptocheirus pinguis
Corophium crassicorne

Species-group E Ampharete arctica

Lumbrineris fragilis
Oioenia fusiformis
Potamilla negleeta

Species-group F Euclymene aollaris

Maldane sarsi
Spiophanes bombyx
Edotea triloba
Duliahia monacantha
Chirodota laevis

Species-group G Cardita bovealis

Asabellides oaulata
Goniada maaulata
Harmothoe imbricata
Ophelina acuminata
Phyllodoce maculata
Diastylis quadrispinosa

99

Species-group H

Amphipholis squamata
Nereis virens
Uniaola irrorata

Species-group I

Yoldia limatula
Aricidea sueaiaa
Eudorella hispida
Erytkrops erythropthalma
Meterythrops robusta

Species-group J

Anemone A

Nassarius trivittatus
Mulinia lateralis
Neomysis ameriaana
Ampelisoa abdita
Melita n. sp.

Species-group K

Eydrobia sp.
Gemma gemma
Tellina agilis

Species-group L

Nemertea D
Nemertea H
Hartmania moorei

100

Species-group M Alvania oavinata

Arioidea quadrilobata
Hhodine loveni
Spio filiaornis
Sternaspis scutata
Leptostylis longimana
Ampelisca agassizi
Anonyx liljeborgi
Metopella angusta

Species-group N Cerebratulus laoteus

Nuoula delphinodonta
Aglaophamus neotenus
Arioidea jeffreysii
Lumbrineris tenuis
Mediomastus ambiseta
Nephtys inaisa
Ninoe nigvipes
Prionospio steenstrupi
Seoloplos sp.
Tharyx sp.
Oligochaeta
Diastylis sculpta
Eudovella trunoatula
Argissa hamatipes

101

vh im

>75

H llllllllllillll

2.50

M KSMWM

2.25

1 W///A

2.10

VL 1 1

<I0

) 6

7

8 9

Fig. 38

CONSTANCY

Constancy of species-groups at site-groups

53
>2

] «l

102

6 7 8 9

FIDELITY

Fig. 39. Fidelity of species-groups to site-groups

103

3 in terms of species-group affiliations but differs in constancy and/or
fidelity levels of species-groups B, C, D and G. Species-groups H, J
and K are most characteristic of site-group 8, as they are present in
medium to high constancy and with high fidelity. Site-group 9 consists
of one station which is qualitatively different from all other stations
in several regards.

Species-group N is unique in that it occurs at all the site-groups
and is present in high to very high constancy at six of the nine
site-groups. Naturally, with such widespread constancy its fidelity to
individual site-groups is very low. Examination of the frequency of
occurrence of the member species of species-group N shows that they
occur at from 53.6 to 87.5% of the stations sampled. This explains the
distinct separation of species-group N from the others in the dendrogram
(Fig. 37). Excepting the special cases of site-groups 6 and 9, we can
characterize the fauna of Casco Bay by species-group N. This group of
very tolerant, numerically dominant species which are undoubtedly
typical of nearshore bottoms over a large area. Superimposed on this
homogeneous fauna are smaller groups of species which are responding to
finer environmental distinctions and hence have a more restricted range
within Casco Bay. It is from among these other groups that initial
changes in community structure, potentially indicative of enviromental
degradation, should be sought.

Several physical and biological parameters are compared in Table 11
on a site-group basis. The observed differences were subjected to
standard analysis of variance and the site-groups differed significantly
(> 95%) from one another in each of the measured parameters. This is
strong evidence that the numerical classification, using only species

104

occurrences and abundances, dissected Casco Bay into ecologically
meaningful components. Analysis of variance also demonstrated that the
site-groups were significantly different in regard to four of the trace
metals, cadmium, chromium, nickel and zinc.

The data was subjected to Duncan's multiple range test to determine
which site-groups differed in the measured parameters. Results of this
procedure are presented in Table 12. Zinc is not included because
Duncan's test is not powerful enough, in this case, to break out the
dissimilar site-groups. Groups represented by the same letter in the
table are not different. For instance, by comparing Tables 11 and 12 we
can conclude that site-group A is significantly deeper than all the
others which do not differ significantly among themselves. Likewise,
site-group 9, located on a mussel reef, has a significantly greater
density than site-groups 1 and 3 which are in turn significantly denser
than the remainder of the groups. Some of the other results are not so
straightforward. In terms of biomass site-groups 1 and 3 are
significantly richer than site-groups 4, 5 and 6, but the intermediate
groups cannot be statistically differentiated from either the high or
low biomass stations.

We are extremely encouraged that the chosen classification
techniques produced groupings that are statistically valid. We are
confident that once all of the data are available we will be able to
provide a comprehensive analysis and benchmark of the present state of
the benthic environment of Casco Bay.

SUMMARY AND TENTATIVE CONCLUSIONS
Casco Bay is a major coastal resource heavily utilized for
commerce, commercial fishing and recreation. Facilities and activities

105

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106

Table 12. Patterns of significant difference between site-groups

based on Duncan's multiple range test. Groups represented
by the same letter (s) are not different.

Site-group
12 3 4 5 6 7 8 9

Depth

A

B

B

B

B

B

B

B

B

Temperature

B

B

B

B

B

B

B

A

A+B

Salinity

A

A+B

A+B

A+B

A+B

B

A+B

C

A+B

Mean Grain

Size

C

A+B+C

C+D

A+B

A

A

D

B+C

C+D

Organic

Carbon

C

A+B

B+C

A+B

A

A

B+C

B+C

A

Density

B

C

B

C

C

C

C

C

A

Biomass

A

A+B

A

B

B

B

A+B

A+B

A+B

Species per

Station

A

C

B

C

D+E

E

C

C+D

C+D+E

Diversity

A+B

A+B

A+B

A+B

B+C

C

A

A+B

A+B+C

Cadmium

B

A

A+B

A

A

A

A+B

B

A

Chromium

B

A+B

B

A

A

A+B

B

B

A+B

Nickel

A+B

A

A+B

A

A

A

B

A+B

A+B

107

potentially threatening to the environment occur throughout the Bay but
are most concentrated in the region of Portland.

Casco Bay is characterized by a boreal climate and a large tidal
range (3 m) . Sediments range from sand in tidally scoured channels to
clay in the inner reaches of the Bay. Interior portions of the Bay have
extremely soft bottom sediments which may be described as fluid mud or
gel. Stations in these areas are occupied by an aberrant community with
low species richness and low density. Further work is needed to fully
document this phenomenon.

Trace metals are not homogeneously distributed throughout Casco
Bay. Sandy and offshore stations tend to be low in metal concentration,
while Portland Harbor appears to contain anthropogenic inputs.
Comparisons with 10 other New England sites confirms that Casco Bay
sediments are impacted in terms of the trace metals sampled.

The fauna of Casco Bay is rich in terms of diversity, density and
biomass. These parameters, and others, are positively correlated with
bottom depth and negatively correlated with mean grain size and organic
carbon content. Most biological parameters are negatively correlated
with at least some of the trace metals. We await the hydrocarbon data
to complete our analysis.

Numerical classifications dissected Casco Bay into nine site-groups
occupied by 14 species-groups. The site-groups are spatially realistic
and differ significantly (>95%) in regard to both physical and
biological factors. One species-group is widely distributed and is
considered typical, boreal shallow-water fauna.

108

ACKNOWLEDGMENTS

This contribution represents the combined efforts of several
persons and organizations. Sampling was facilitated by the excellent
support of Paul DeRocher and Mike Dunton aboard the Maine Department of
Marine Resources (DMR) vessel R/V Explorer. Joslyn Kelly and Robert
Wells of the Bigelow Laboratory, and Ellen Montgomery of Colby College
were responsible for much of the pre-sorting of faunal samples. Trace
metal analyses were accomplished by Vincent Zdanowicz of the National
Marine Fisheries Service (NMFS) . Advice on the interpretation of the
metals data was provided by Henri Gaudette and Berry Lyons of the
University of New Hampshire. David Sampson of DMR and Jean Garside of
the Bigelow Laboratory provided statistical advice and data processing
support. Drafting was undertaken by Jim Rollins of the Bigelow
Laboratory. Special thanks go to Margaret Colby, Pat Oathout and Teve
MacFarland for the painstaking manuscript typing, and to Robert Reid of
NMFS for general support and encouragement throughout.

109

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Parker, C, and N. Garfield, III. 1981b. Station data obtained in

Northwestern Gulf of Maine coastal area, October 1979. 117 pp.
Pielou, E.C. 1970. An introduction to mathematical ecology. Wiley-

Interscience. New York. 286 pp.

112

Rosenberg, R. 1973. Succession in benthic macrofauna in a Swedish
fjord subsequent to the closure of a sulphite pulp mill. Oikos
24: 244-258.

Rowe, G.T., P.T. Polloni and J.I. Rowe. 1972. Benthic community para-
meters in the lower Mystic River. Int. Rev. ges . Hydrobiol. 57:
573-584.

Shorey, W.K. 1973. Macrobenthic ecology of a sawdust-bearing substrate
in the Penobscot River estuary (Maine). J. Fish. Res. Bd. Canada
30: 493-497.

U.S. Fish and Wildlife Service. 1980. An ecological characterization
of coastal Maine. FWS/OBS-80129. Department of Interior,
Washington, D.C.

Verrill, A.E. 1874. Explorations of Casco Bay in 1873. Proc. Amer.
Assoc. Adv. Sci. (Portland Meeting) 22: 340-395.

1 13

Appe nd ix 1 .
Sediment data by station

114

Table A-l . The percentage of sand, silt and clay particles in each
Casco Bay sediment sample taken in April 1980.

Station % sand % silt % clay

2

53.7

28.0

18.3

3

58.6

19.5

22.0

4

64.0

17.4

18.6

5

32.0

34.5

33.5

6

15.3

47.9

36.8

7

9.1

46.5

44.4

8

63.3

21.3

15.4

9

99.2

.3

.4

10

20.8

29.0

50.2

11

34.6

46.4

19.0

12

36.6

34.6

28.9

13

35.1

35.0

30.0

15

20.4

45.6

34.0

16

4.9

48.8

46.3

17

1.4

49.2

49.4

18

71.5

13.7

14.8

19

21.7

40.1

38.1

20

16.6

44.3

39.1

21

10.5

45.5

44.0

22

89.9

5.2

4.9

23

30.5

40.2

29.4

24

10.0

54.4

35.6

25

13.0

38.8

48.2

26

3.1

53.8

43.0

27

7.1

48.6

44.4

28

26.7

40.3

33.0

29

23.0

42.7

34.4

30

6.2

40.1

53.7

31

1.7

48.0

50.3

32

4.4

47.8

47.8

33

14.7

47.9

37.4

115

Station

% san

34

3.0

35

1.7

36

15.2

37

45.7

38

1.9

39

1.7

40

2.1

41

20.7

42

7.3

43

2.4

44

16.5

45

6.1

46

57.2

47

65.5

48

73.7

49

50

97.2

51

41.9

52

19.7

53

69.6

54

66.4

55

73.5

56

27.8

57

9.3

58

5.3

% silt

49.4
50.8
39.9
33.6
47.3
50.4
46.4
47.9
51.9
50.8
50.6
52.2
23.9
21.3
16.1

1.2
31.3
42.4
18.8
18.6
14.3
39.4
48.4
54.6

%clay

47.6
47.5
44.9
20.7
50.9
48.0
51.5
31.4
40.9
46.8
32.9
41.7
19.0
13.2
10.2

1.5

26.8
37.9
11.6
15.0
12.2
32.7
42.3
40.0

117

Appendix 2 .
Faunal data by station

118

CRUISE EX8001

STATION 02

RANK
1
2
3
4
5
6
7
B
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59

SPECIES NAME
AMPELISCA AGASSIZI
PRIONOSF'IO STEENSTRURI
NINOE N1GRIPES
CASCO BIGELOUI
MEDIOMASTUS AMBISETA
PHOT IS MACROCOXA
THARYX SP.
AMFHARETE ARCTICA
SCOLOPLOS SP.
HARPINIA F'ROPINOUA
ASABELLIDES OCULATA
CRENELLA [lECUSSATA
RHODINE LOVENI
SABELLA PENICILLUS
EUCIORELLA TRUNCATULA
IiIASTYLIS SCULPTA
CEREBRATULUS LACTEUS
PERIPLOMA PAPYRATIUM
LUMBRINERIS FRAC-TLIS
GONIADA MACULATA
PARAONIS GRACILIS
STENOPLEUSTES INERMIS
PHOLOE MINUTA
SPIO FILICORNIS
PETALQSARSIA DECLIVIS
MYA ARENARIA
AMFHARETE ACUTIFRONS
ARICIDEA SUECICA
MELLINA CRISTATA
LEPTOSTYLIS LONGIMANA
ERICTHONIUS RUBRICORNIS
NEMERTEA D
NEMERTEA C
BRADA GRANOSA
LUMBRINERIS TENUIS
NEPHTYS INCISA
HARMOTHOE IMBRICATA
STERNASPIS SCUTATA
PRAXILLELLA GRACILIS
OPHELINA ACUMINATA
[IIASTYLIS ABBREVIATA
HIASTYLIS GUADRISPINOSA
EDOTEA TRILOBA
ARGISSA HAMATIPES
THYASIRA FLEXUOSA
NUCULA ANNULATA
PITAR MORRHUANA
EXOGONE VERUGA
PHYLLOIiOCE MUCOSA
OLIGOCHAETA
LA0N1CE CIRRATA
MAYERELLA LIMICOLA
CAMPYLASFIS RUBICUNPA
DIASTYLIS CORNUIFER
OXYUROSTYLIS SMITHI
DULICHIA MONACANTHA
PHOXOCEPHALUS HOLBOLLI
PROTOMEDEIA FASCIATA
MONOCULODES N.SP.

COUNT

CUM COUNT

V.

CUM X

1793.

1793.

68. 10

68.10

456.

2249.

17.32

85.42

44.

2293.

1.67

87.09

42.

2335.

1 .60

88.68

26.

2361 .

0.99

89.67

26.

2387.

0.99

90.66

24.

2411 .

0.91

91 .57

21

2432.

0.80

92.37

17

2449.

0.65

93.01

17

2466.

0.65

93.66

14

2480.

0.53

94.19

12

2492.

0.46

94.64

12

2504.

0.46

95.10

10

2514.

0.3B

95.48

7

n 521

0.27

95.75

7

2528.

0.27

96.01

6

2534.

0.23

96.24

6

2540.

0.23

96.47

6

2546.

0.23

96.70

6

2552.

0.23

96.92

5

2557.

0. 19

97.11

5

^*562 .

0.19

97.30

4

2566.

0.15

97.46

4

2570.

0.15

97.61

4

2574.

0. 15

97.76

3

2577.

0.11

97.87

3

2580.

0.11

97.99

3

2583.

0.11

98.10

3

2586.

0.11

98.21

3

2589 .

0.11

98.33

3

2592.

0.11

98.44

2

2594.

0.08

98.52

2

, 2596.

0.08

98.59

2

2598.

0.08

98.67

2

2600.

0.08

98.75

2

2602.

0.08

98.82

'y

2604.

0.08

98.90

2

2606.

0.08

98.97

r>

2608.

0.08

99.05

2

2610.

O.OB

99.13

2

2612.

0.08

99.20

■y

, 2614.

0.08

99.28

*?

2616.

0.08

99.35

2

, 2618.

0.08

99.43

2619.

0.04

99.47

2620.

0.04

99.51

2621 .

0.04

99.54

2622.

0.04

99.58

2623.

0.04

99.62

2624.

0.04

99.66

2625.

0.04

99.70

. 2626.

0.04

99.73

2627.

0.04

99.77

2628.

0.04

99.81

2629.

0.04

99.85

2630.

0.04

99.89

2631.

0.04

99.92

2632.

0.04

99.96

2633.

0.04

100.00

NUMBER OF SPECIES 59

NUMBER OF INDIVIDUALS 2633.
INDIVIDUALS PER M2 26330

119

CRUISE EX8001

STATION 03

RANK SPECIES NAME

1 AMPELISCA AGASSIZI

2 HARLOORS TUBICOLA

3 PRIONOSPIO STEENSTRURI

4 RHODINE LOOENI

5 MALDANE SARSI

6 PROTOMEDEIA FASCIATA

7 CRENELLA DECUSSATA

8 LEPTOCHEIRUS PINGUIS

9 THARYX SP.

10 ERICTHONIUS RUBRICORNIS

11 HARRINIA PROPINQUA

12 MASTYLIS 0UAPRISP1N0SA

13 AMPHARETE ARCTICA

14 EUDORELLA TRUNCATULA

15 PHYLLODOCE MUCOSA

16 THYASIRA FLEXUOSA

17 AMPELISCA MACROCEPHALA

18 ASABELLIDES OCULATA

19 PHOT IS MACROCOXA

20 TEREBELLID B

21 MEDIOMASTUS AMBISETA

22 OPHELINA ACUMINATA

23 NINOE N I GRIPES

24 SPIO FILICORNIS

25 ASTARTE UNDATA

26 NUCULA DELPHINOBONTA

27 LUMBRINERIS FRAGILIS

28 AMPHARETE ACUTIFRONS

29 PERIPLOMA PAPYRATIUM

30 UNCIOLA IRRORATA

31 SABELLA PENICILLUS

32 PHOLOE MINUTA

33 DIASTYLIS ABBREVIATA

34 DIASTYLIS SCULPTA

35 EBOTEA TRILOBA

36 STEREOBALANUS CANADENSIS

37 CERIANTHUS BOREALIS

38 AMPHIPHOLIS SOUAMATA

39 HARTMANIA MOOREI

40 HARMOTHOE IMBRICATA

41 LEPTOSTYLIS LONGTMANA

42 MUNNA FABRICII

43 ANONYX LILJEBORGI

44 STENOPLEUSTES INERMIS

45 NOTOPLANA ATOMATA

46 PERIPLOMA LEANUM

47 CARDITA BOREALIS

48 OUENIA FUSIFORMIS

49 SYLLIS GRACILIS

50 EUCLYMENE COLLARIS

51 PHYLLODOCE MACULATA

52 ETEONE LONGA

53 MELINNA CRISTATA

54 NEPHTYS INCISA

55 PRAXILLELLA GRACILIS

56 STERNASPIS SCUTATA

57 GITANOPSIS SP.

58 HALIMEHON SP .

59 DULICHIA MONOCAHTHA

60 NEMERTEA G

61 THRACIA CONRADI

62 CHLAMYS ISLANIUCA

63 LYONSIA HYALINA

64 MODIOLUS MODIOLUS

65 TRICHOBRANCHUS GLACIAL IS

66 TROCHOCHAETA MULTISETOSA

67 EXOGONE HEBES

68 PHERUSA AFFINIS

69 AGLAOPHAMUS CIRCINATA

70 SCALIBREGMA INFLATUM

71 SCOLOPLOS SP.

72 GONIADA MACULATA

73 ARICIDEA JEFFREYSII

74 PARAONIS GRACILIS

75 TEREBELLIDES STROEMI

76 METERYTHROPS ROBUSTA

77 PETALOSARSIA DECLIVIS

78 DIASTYLIS CORNUIFER

79 CANCER BOREALIS

80 PLEUSYMTES GLABER

81 BATHYMEDON SP .

82 PHOXOCEPHALUS HOLBOLLI

83 ARGISSA HAMATIPES

84 PONTOGENEIA INERMIS

85 COROPHIUM CRASSICORNE

86 METOPELLA ANGUSTA

C0UN1

run COUNT

V.

CUM X

385.

385.

27.66

27.66

217

602.

15.59

43.25

134

736.

9.63

52.87

67.

803.

4.81

57.69

65.

868.

4.67

62.36

47.

915.

3.38

65.73

45.

960.

3.23

68.97

40.

1000.

2.87

71 .84

39.

1039.

2.80

74.64

36

1075.

2.59

77.23

23

1098.

1 .65

78.88

21

1119.

1 .51

80.39

18

1137.

1 .29

81 .68

18

1155.

1.29

82.97

15

1170.

1 .08

84.05

11

1181 .

0.79

84.84

11

1192.

0.79

85.63

10

1202.

0.72

86.35

10

1212.

0.72

87.07

9

1221.

0.65

87.72

9

1230.

0.65

88.36

9

1239.

0.65

89.01

8

1247.

0.57

89.58

7

1254.

0.50

90.09

6

1260.

0.43

90.52

6

1266.

0.43

90.95

6

1272.

0.43

91 .38

6

1278.

0.43

91 .81

5

1283.

0.36

92.17

5

1288.

0.36

92.53

4

1292.

0.29

92.82

4

1296.

0.29

93.10

4

1300.

0.29

93.39

4

1304.

0.29

93.68

4

1308.

0.29

93.97

3

1311 .

0.22

94.18

3

1314.

0.22

94.40

3

1317.

0.22

94.61

3

1320.

0.22

94.83

3

1323.

0.22

95.04

3

1326.

0.22

95.26

3

1329.

0.22

95.47

3

1332.

0.22

95.69

3

1335.

0.22

95.90

2

1337.

0.14

96.05

2

1339.

0.14

96.19

•3

1341 .

0.14

96.34

2

1343.

0.14

96.48

2

1345.

0.14

96.62

2

1347.

0.14

96.77

-»

1349.

0.14

96.91

o

1351.

0.14

97.05

2

, 1353.

0.14

97.20

2

1355.

0.14

97.34

2

1357.

0.14

97.49

2

1 359 .

0.14

97.63

i

1361.

0.14

97.77

2

1363.

0. 14

97.92

2

1365.

0.14

98.06

1366.

0.07

98.13

1367.

0.07

98.20

1368.

0.07

98.28

1369.

0.07

98.35

1370.

0.07

98.42

1371.

0.07

98.49

1372.

0.07

98.56

1373.

0.07

98.63

1374.

0.07

98.71

1375.

0.07

98.78

1376.

0.07

98.85

1377.

0.07

98.92

1378.

0.07

98.99

1379.

0.07

99.07

1380.

0.07

99.14

1381.

0.07

99.21

1382.

0.07

99.28

1383.

0.07

99.35

1384.

0.07

99.42

1 385 .

0.07

99.50

1386.

0.07

99.57

1387.

0.07

99.64

1388.

0.07

99.71

1389.

0.07

99.78

1390.

0.07

99.86

1391.

0.07

99.93

1392.

0.07

100.00

NUMBER OF SPECIES 86

NUMBER OF INDIVIDUALS 1392.
INDIVIDUALS PER M2 13920

120

CRUISE EX8001

STATION 04

GRAB 1

RANK
1

3

4

5

6

7

8

9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72

SPECIES NAME
PRIONOSPIO STEENSTRUPI
AMPELISCA AGASSIZI
HAPLOOPS TUBICOLA
CRENELLA DECUSSATA
NUCULA DELPHINOPONTA
MEEHOMASTUS AMBISETA
PHOXOCEPHALUS HOLBOLLI
HARPINIA PROPINQUA
EUDORELLA TRUNCATULA
SABELLA PEHICILLUS
PHOT IS MACROCOXA
LEPTOCHEIRUS PINGUIS
PHYLLOIiOCE MUCOSA
DULICHIA MONOCANTHA
AMPELISCA MACROCEPHALA
ASTARTE BOREALIS
AMPHARETE ARCTICA
NINOE NIGRIPES
PERIPLOMA PAPYRATIUM
THARYX SP.

DIASTYLIS QUADRISPINOSA
ORCHOMENELLA PINGUIS
ETEONE LONGA
DIASTYLIS SCULPTA
HARMOTHOE IMBRICATA
CASCO BIGELOWI
TEREBELLIDAE
RHOHINE LOVENI
GONIADA MACULATA
STENOPLEUSTES INERMIS
ERICTHONIUS RUBRICORNIS
NUCULA ANNULATA
ALVANIA CARINATA
POLYDORA OUAPRILOBATA
PHOLOE MINUTA
LUMBRINERIS FRAGILIS
CERASTODERMA PINNULATUM
THYASIRA FLEXUOSA
MELINNA CRISTATA
EDOTEA TRILOBA
COROPHIUM CRASSICORNE
ANONYX LILJEBORGI
CARDITA BOREALIS
AMPHARETE ACUTIFRONS
TRICHOBRANCHUS GLACIALIS
ASABELLIDES OCULATA
SCOLOPLOS SP.
OPHELINA ACUMINATA
ARICIPEA JEFFREYSII
NEPHTYS INCISA
MUNNA FABRICII
CHIRIDOTA LAEVIS
NEMERTEA G
CERIANTHUS BOREALIS
OENOPOTA BICARINATA
MYA ARENARIA
LACUNA VINCTA
TEREBELLIDES STROEMI
NEREIS GRAY I
OLIGOCHAFTA
SPIO FILICORNIS
LAONICE CIRRATA
OUIENIA FUSIFORMIS
STAURONEREIS CAECUS
PARAONIS GRACILIS
MALDANE SARSI
BRADA UILLOSA
PHERUSA AFFINIS
HALIMEDON SP.
ARGISSA HAMATIPES
CEREBRATULUS LACTEUS
PHYLUM A

COUNT CUM COUNT

V.

CUM X

1215

1215.

41 .48

41 .48

1020

2235.

34.82

76.31

137

2372.

4.68

80.98

79

2451 .

2.70

83.68

48

2499.

1 .64

85.32

46

2545.

1.57

86.89

34

2579.

1.16

88.05

33

2612.

1 .13

89. 18

29

2641 .

0.99

90.17

19

2660.

0.65

90.82

17

2677.

0.58

91 .40

16

2693.

0.55

91 .94

15

2708.

0.51

92.45

15

2723.

0.51

92.97

14

2737.

0.48

93.44

10

2747.

0.34

93.79

10

2757.

0.34

94.13

9

2766.

0.31

94.43

8

2774.

0.27

94.71

8

2782.

0.27

94.98

8

2790.

0.27

95.25

8

2798.

0.27

95.53

7

2805.

0.24

95.77

7

2812.

0.24

96.01

6

2818.

0.20

96.21

6

2824.

0.20

96.41

5

2829.

0.17

96.59

5

2834.

0.17

96.76

5

2839.

0.17

96.93

5

2844.

0.17

97.10

5

2849.

0. 17

97.27

4

2853.

0.14

97.40

4

2857.

0.14

97.54

4

2861 .

0.14

97.68

4

2865.

0.14

97.81

4

2869.

0.14

97.95

3

2872.

0. 10

98.05

3

2875.

0.10

98. 16

3

2878.

0.10

98.26

3

2881 .

0.10

98.36

3

2884.

0.10

98.46

3

2887.

0.10

98.57

2

2889.

0.07

98.63

2

2891 .

0.07

98.70

2

2893.

0.07

98.77

2

2895.

0.07

98.84

2

2897.

0.07

98.91

2

2899.

0.07

98.98

2

2901.

0.07

99.04

2

2903.

0.07

99.11

n

2905.

0.07

99.18

-y

. 2907.

0.07

99.25

o

2909.

0.07

99.32

2

2911 .

0.07

99.38

2912.

0.03

99.42

2913.

0.03

99.45

2914.

0.03

99.49

2915.

0.03

99.52

2916.

0.03

99.56

2917.

0.03

99.59

2918.

0.03

99.62

2919.

0.03

99.66

2920.

0.03

99.69

2921 .

0.03

99.73

2922 .

0.03

99.76

2923.

0.03

99.79

2924.

0.03

99.83

2925.

0.03

99.86

2926.

0.03

99.90

2927.

0.03

99.93

2928.

0.03

99.97

2929.

0.03

100.00

NUMBER OF SPECIES 72

NUMBfR OF INDIVIDUALS 2929.
INDIVIDUALS PER M2 29290

121

CRUISE EX8001

STATION 05

RANK SPECIES NAME

1 AMFELISCA AGASSIZI

2 MALDANE SARSI

3 HAPLOOPS TUBICOLA

4 PRIOHOSPIO STEENSTRUFI

5 AMPHARETE ARCTICA

6 RHOniNE LOVENI

7 THARYX SP.

8 SPIO FILICORNIS

9 ASABELLIDES OCULATA

10 LUMBRINERIS FRAGILIS

11 ERICTHONIUS RUBRICORNIS

12 BYBLIS GAIMARIlI

13 MEDIOMASTUS AMBISETA

14 CAPRELLA UNICA

15 ASTARTE UNDATA

16 NINOE NIGRIPES

17 DIASTYLIS OUADRISPINOSA

18 MELINNA CRISTATA

19 HARPINIA PROPINOUA

20 PHOLOE MINUTA

21 SABELLA PENICILLUS

22 CARBITA BOREALIS

23 STERNASPIS SCUTATA

24 EUDORELLA TRUNCATULA

25 PERIPLOMA PAPYRATIUM

26 NUCULA DELFHINODONTA

27 CEREBRATULUS LACTEUS

28 PHYLLODOCE MUCOSA

29 CRENELLA DECUSSATA

30 THYASIRA FLEXUOSA

31 GONIAHA MACULATA

32 BULICHIA MOHACANTHA

33 TRICHOBRANCHUS GLACIALIS

34 PARAONIS GRACILIS

35 NOTOMASTUS LATERICUS

36 BIF'LOCIRRUS HIRSUTUS

37 AEGININA LONGICORNIS

38 PHOT IS MACROCOXA

39 LEPTOCHEIRUS PINGUIS

40 HALIMEDON SP.

41 ALVANIA CARINATA

42 MODIOLUS MODIOLUS

43 AMPHIPHOLIS SOUAMATA

44 OLIGOCHAETA

45 STAURONEREIS CAECUS

46 NEPHTYS INCISA

47 SCALIBREGMA INFLATUM

48 LAONICE CIRRATA

49 LEPTOSTYLIS LONGIMANA

50 PTILANTHURA TENUIS

51 MUNNA FABRICII

52 ORCHOMENELLA PINGUIS

53 PHOXOCEPHALUS HOLBOLLI

54 UNCIOLA IRRORATA

55 PLEUSTES PANOPLUS

56 METOPELLA ANGUSTA

57 PONTOGENEIA INERMIS

58 ANONYX LILJEBORGI

59 AMPELISCA MACROCEPHALA

60 PHASCOLION STROMRI

61 CERIANTHUS BOREALIS

62 NUCULA ANNULATA

63 CERASTODERMA PINNULA1UM

64 CHIRIBOTA LAEVIS

65 STRONGYLOCENTROTUS DROEBACHIENSIS

66 OPHIOPHOLIS ACULEATA

67 OPHIURA SARSI

68 PARAF'IONOSYLLIS LONGOCIRRATA

69 SFHAERODOROPSIS MINUTA

70 EUSYLLIS BLOMSTRAHDI

71 ARICIBEA OUADRILOBATA

72 PRAXILLELLA PRAETERMISSA

73 ETEONE LONGA

74 POTAMILLA MEGLECTA

75 PHYLLOBOCE MACULATA

COUNT

CUM COUNT

■/.

CUM X

1115.

1115.

42.12

42.12

J80.

1 495 .

14.36

56.48

311 .

1806.

11 .75

68.23

183.

1989.

6.91

75.14

162.

2151 .

6. 12

81 .26

130.

2281 .

4.91

86.17

43.

2324.

1.62

87.80

26.

2350.

0.98

88.78

24

2374.

0.91

89.69

20

2394.

0.76

90.44

18

2412.

0.68

91.12

16

2428.

0.60

91 .73

16

2444.

0.60

92.33

11

2455.

0.42

92.75

11 .

2466.

0.42

93.16

11

2477.

0.42

93.58

10

2487.

0.38

93.96

10

2497.

0.38

94.33

9

2506.

0.34

94.67

9

2515.

0.34

95.01

9

2524.

0.34

95.35

8

2532.

0.30

95.66

8

2540.

0.30

95.96

7

2547.

0.26

96.22

6

2553.

0.23

96.45

6

2559.

0.23

96.68

5

2564.

0.19

96.86

5

2569.

0.19

97.05

4

2573.

0.15

97.20

4

2577.

0.15

97.36

4

2581 .

0. 15

97.51

3

2584.

0.11

97.62

3

2587.

0.11

97.73

3

2590.

0.11

97.85

3

2593.

0.11

97.96

3

2596.

0.11

98.07

2

2598.

0.08

98.15

2

2600.

0.08

98.22

i

2602.

0.08

98.30

r>

2604.

0.08

98.38

2

2606.

0.08

98.45

T

2608.

0.08

98.53

"1

2610.

0.08

98.60

2

2612.

0.08

98.68

O

2614.

0.08

98.75

•?

2616.

0.08

98.83

2

2618.

0.08

98.90

2

2620.

0.08

98.98

2621.

0.04

99.02

■ 2622.

0.04

99.06

2623.

0.04

99.09

2624.

0.04

99.13

2625.

0.04

99.17

2626.

0.04

99.21

2627.

0.04

99.24

2628.

0.04

99.28

2629.

0.04

99.32

2630.

0.04

99.36

2631 .

0.04

99.39

2632.

0.04

99.43

2633.

0.04

99.47

2634.

0.04

99.51

2635.

0.04

99.55

2636.

0.04

99.58

2637.

0.04

99.62

2638.

0.04

99.66

2639.

0.04

99.70

2640.

0.04

99.73

2641.

0.04

99.77

2642.

0.04

99.81

2643.

0.04

99.85

2644.

0.04

99.89

2645.

0.04

99.92

2646.

0.04

99.96

2647.

0.04

100.00

NUMBER OF SPECIES 75

NUMBER OF - INDIVIDUALS 2647.
INDIVIDUALS PER M2 26470

122

CRUISE EXB001

STATION 06

RANN SPECIES NAME

1 PRIONOSFIO STEENSTRUPI

2 STERNASPIS SCUTATA

3 MEDIOMASTUS AMBISETA

4 THARYX SP.

5 SPIO FILICORNIS

6 NINOE NIGRIPES

7 AMPHIPHOLIS SQUAMATA

8 HALIMEUON SP.

9 CEREBRATULUS LACTEUS

10 EUDORELLA TRUNCATULA

11 OUENIA FUSIFORMIS

12 PAR AON IS GRACILIS

13 STENOPLEUSTES INERMIS

14 SCOLOPLOS SP.

15 AGLAOPHAMUS NEOTENUS

16 ARICIHEA QUAHRILOBATA

17 NUCULA BELPHINOBONTA
IB ALVANIA CARINATA

19 OLIGOCHAETA

20 MALDANE SARSI

21 ARICIHEA JEFFREYSII

22 AMPHARETE ARCTICA

23 DIPLOCIRRUS HIRSUTUS

24 LUMBRINERIS FRAGILIS

25 STEREOBALANUS CANADENSIS

26 HULICHIA MONOCANTHA

27 EDOTEA TRILOBA

28 HARTMANIA MOOREI

29 ARGISSA HAMATIPES

30 MONOCULODES TESSELATUS

31 APISTOBRANCHUS TULLBERGI

32 NEMERTEA D

33 BATHYMEDON SP.

34 HARPINIA PROPINGUA

35 METOPELLA ANGUSTA

36 AMPELISCA AGASSIZI

37 MELITA N.SP.

38 SABELLA PENICILLUS

39 THYASIRA FLEXUOSA

40 PERIPLOMA PAPYRATIUM

41 ANEMONE A

42 ECHIURUS ECHIURUS

43 NEMERTEA C

44 MAYERELLA LIMICOLA

45 ORCHOMENELLA PINGUIS

46 PHOT IS MACROCOXA

47 TEREBELLIBAE

48 PRAXILLELLA GRACILIS

49 TRICHOBRANCHUS GLACIALIS

50 NEPHTYS INCISA

51 RHODINE LOVENI

52 ARICIHEA SUECICA

53 PHOLOE MINUTA

54 SPIOPHANES BOMBYX

55 PRAXILLELLA SP .

56 CERASTOBERMA PINNULATUM

COUNT CUM COUNT

7.

CUM 7.

371

371.

56 . 55

56.55

33

404.

5.03

61 .59

30

434.

4.57

66. 16

21

455.

3.20

69.36

16

471.

2.44

71 .80

12

483.

1 .83

73.63

11

494.

1 .68

75.30

9

503.

1.37

76.68

7

510.

1.07

77.74

7

517.

1.07

78.81

7

524.

1.07

79.88

7

531.

1.07

80.94

6

537.

0.91

81 .86

6

543.

0.91

82.77

6

549.

0.91

83.69

6

j jo .

0.91

84.60

6

561.

0.91

85 . 52

6

567.

0.91

86.43

•j

572.

0.76

87.19

5

577.

0.76

87.96

5

582.

0.76

88.72

5

587.

0.76

89.48

5

592.

0.76

90.24

5

597.

0.76

91.01

4

601.

0.61

91.62

4

605.

0.61

92.23

4

609.

0.61

92.84

4

613.

0.61

93.44

3

616.

0.46

93.90

3

619.

0.46

94.36

3

622.

0.46

94.82

2

624.

0.30

95.12

2

626.

0.30

95.43

2

628.

0.30

95.73

2

630.

0.30

96.04

3

632.

0.30

96.34

2

634.

0.30

96.65

-»

636.

0.30

96.95

->

638.

0.30

97.26

2

640.

0.30

97.56

641 .

0.15

97.71

642.

0.15

97.87

643.

0. 15

98.02

644.

0.15

98.17

645.

0.15

98.32

646.

0. 15

98.48

647.

0.15

98.63

648.

0.15

98.78

649.

0. 15

98.93

650.

0. 15

99.08

651.

0.15

99.24

652.

0. 15

99.39

653.

0. 15

99.54

654.

0.15

99.69

655.

0.15

99.85

656.

0. 15

100.00

NUMBER OF SPECIES 56

NUMBER OF INDIVIDUALS 656.

INIiIUIIiUALS PER M2 6560

CRUISE EXB001

STATION 07

RANK
1

7
S
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24

SPECIES NAME
EUDORELLA TRUNCATULA
PRIONOSFIO STEENSTRUPI
MASTYLIS SCULPTA
AGLAOPHAMUS NEOTENUS
NEPHTYS INCISA
ERYTHROPS ERYTHROPHTHALMA
ARICIHEA SUECICA
MEDIOMASTUS AMSISETA
CERE&RATULUS LACTEUS
SCOLOPLOS SP.
EUDORELLA HISPIDA
HALIMEDON SP.
LUMBRINERIS FRAGILIS
THARYX SF'.
DULICHIA MONOCANTHA
ORCHOMENELLA FINGUIS
PATHYMEHON SP .
NEOMYSIS AMERICANA
GEMMA GEMMA

CERASTODERMA PINNULATUM
NASSARIUS TRIVITTATUS
ARICIHEA JEFFREYSII
ETEONE LONGA
ARGISSA HAMATIPES

IJUMEiER OF SPECIES

24

NUMPER Of INMUIHUALS 732.

INDIVIDUALS PER M2 7320

123

COUNT CUM COUNT

X

CUM '/.

491

491 .

67.08

67.08

87

57B.

11 .89

78.96

51

629.

6.97

85.93

37

666.

5.05

90.98

11

677.

1.50

92.49

11

688.

1.50

93.99

8

696.

1.09

95.08

5

701.

0.68

95.76

4

705.

0.55

96.31

3

708.

0.41

96.72

3

711 .

0.41

97. 13

3

714.

0.41

97.54

2

716.

0.27

97.81

2

718.

0.27

98.09

2

720.

0.27

98.36

2

722.

0.27

98.63

2

724.

0.27

98.91

2

726.

0.27

99.18

727.

0. 14

99.32

728.

0. 14

99.45

729.

0.14

99.59

730.

0. 14

99.73

731.

0. 14

99.86

732.

0.14

100.00

CRUISE EX8001

STATION 08

RANK SPECIES NAME

1 PRIONOSPID STEENSTRUFI

2 MEDIOMASTUS AMBISETA

3 EUDORELLA TRUNCATULA

4 LUMBRINERIS TENUIS

5 ARICHiEA JEFFREYSII

6 NINOE NIGRIPES

7 APISTOBRANCHUS TULLBERGI

8 NUCULA DELPHINODONTA

9 ARGISSA HAMATIPES

10 OLIGOCHAETA

11 5COLOPL0S SP.

12 AGLAOPHAMUS NEOTENUS

13 PHYLLODOCE MUCOSA

14 OUENIA FUSIFORMIS

15 STERNASPIS SCUTATA

16 PARAONIS GRACILIS

17 ETEONE LONGA

18 CRENELLA DECUSSATA
1? ORCHOMENELLA PINGUIS

20 PHOLOE MINUTA

21 AMPHARETE ARCTICA

22 OPHELINA ACUMINATA

23 NEMERTEA [r

24 CHIRIDOTA LAEVIS

25 PHERUSA AFFINIS

26 THARYX SP.

27 LUMBRINERIS FRAGILIS

28 NEOMYSIS AMERICANA

29 AMPELISCA VADORUM

30 CASCO BIGELOUI

31 CYLICHNA GOULIU

32 THYASIRA FLEXUOSA

33 GEMMA GEMMA

34 YOLDIA L1MATULA

35 PERIPLOMA PAPYRATIUM

36 NEMERTEA C

37 CERIANTHUS BOREALIS

38 SABELLA PENICILLUS

39 HARTMANIA MOOREI

40 ETEONE FLAVA

41 METOPELLA ANGUSTA

NUMBER OF SPECIES 41

NUMBCH OF INDIVIDUALS 1304.
INDIVIDUALS PER M2 13040

124

COUNT CUM COUNT

V.

CUM X

977

977.

74.92

74.92

56

1033.

4.29

79.22

42

1075.

3.22

82.44

37

1112.

2.84

85.28

18

1130.

1 .38

86.66

16

1146.

1 .23

87.88

15

1161 .

1.15

89.03

13

1174.

1 .00

90.03

13

1187.

1.00

91.03

11

1198.

0.84

91 .87

11

1209.

0.84

92.71

11

1220.

0.84

93.56

10

1230.

0.77

94.33

8

1238.

0.61

94.94

6

1244.

0.46

95.40

6

1250.

0.46

95.86

6

1256.

0.46

96.32

5

1261 .

0.38

96.70

5

1266.

0.38

97.09

4

1270.

0.31

97.39

4

1274.

0.31

97.70

3

1277.

0.23

97.93

2

1279.

0. 15

98.08

.■>

1281 .

0.15

98.24

T

1283.

0. 15

98.39

T

1285.

0.15

98.54

2

1287.

0.15

98.70

-y

1289.

0.15

98.85

2

1291.

0.15

99.00

o

1293.

0. 15

99. 16

1294.

0.08

99.23

1295.

0.08

99.31

1296.

0.08

99.39

1297.

0.08

99.46

1298.

0.08

99.54

1299.

0.08

99.62

1300.

0.08

99.69

1301 .

0.08

99.77

1302.

0.08

99.85

1303.

0.08

99.92

1304.

0.08

100.00

CRUISE EX8001

STATION 09

RANK SPECIES NAME

1 OLIGOCHAETA

2 ARCHIANNELIDA

3 PARAONIS LYRA

4 THARYX SP.

5 CEREBRATULUS LACTEUS

6 EXOGONE VERUGA

7 PRAXILLELLA PRAETERMISSA

8 EXOGONE HEBES

? ARICIDEA JEFFREYSII

10 AMPHARETE ARCTICA

11 CHIRODOTEA COECA

12 STAURONEREIS RUDOLPHI

13 SYLLIS CORNUTA

14 LUMBRINERIS ACUTA

15 POLYCIRRUS PHOSPHOREUS

16 PLATYHELMINTHES

17 PRIONOSPIO STEENSTRUPI

18 PHOLOE MINUTA

19 LUMBRINERIS TENUIS

20 ASTARTE BOREALIS

21 NASSARIUS TRIVITTATUS

22 LUMBRINERIS FRAGILIS

23 CERASTOIIERMA PINNULATUM

24 OUENIA FUSIFORMIS

25 POTAMILLA NEGLECTA

26 OPHIOGLYCERA GIGANTEA

27 GONIADA MACULATA

28 AGLAOPHAMUS CIRCINATA

29 DIASTYLIS SCULPTA

30 PSAMMONYX NOBILIS

NUMBER OF SPECIES

30

NUMBER OF INDIVIDUALS 1591.
INDIVIDUALS PER M2 15910

125

COUNT CUM COUNT

%

CUM '/.

776

776.

48.77

48.77

186

962.

11 .69

60.47

161

1123.

10.12

70.58

95

1218.

5.97

76.56

72

1290.

4.53

81 .08

63

1353.

3.96

85.04

48

1401.

3.02

88.06

37

1438.

2.33

90.38

33

1471.

2.07

92.46

25

1496.

1.57

94.03

22

1518.

1.38

95.41

19

1537.

1.19

96.61

10

1547.

0.63

97.23

9

1556.

0.57

97.80

7

1563.

0.44

98.24

4

1567.

0.25

98.49

4

1571 .

0.25

98.74

3

1574.

0. 19

98.93

3

1577.

0.19

99.12

2

1579.

0.13

99.25

2

1581.

0.13

99.37

2

1583.

0. 13

99.50

1584.

0.06

99.56

1585 •

0.06

99.62

1586.

0.06

99.69

1587.

0.06

99.75

1588.

0.06

99.81

1589.

0.06

99.87

1590.

0.06

99.94

1591 .

0.06

100.00

126

CRUISE EX8001

STATION 10

GRAB 1

RANK SPECIES NAME

1 PRI0N0SPI0 STEENSTRUPI

2 NUCULA BELPHINODONTA

3 SPIO FILICORNIS

4 AMPELISCA AGASSIZI

5 THARYX SP.

6 ARCTICA ISLANBICA

7 AMPHARETE ACUTIFRONS

8 CASCO BIGELOUI

9 NINOE NI GRIPES

10 MEIHOMASTUS AMBISETA

11 EDOTEA TRILOBA

12 CRENELLA BECUSSATA

13 MALBANE SARSI

14 STERNASPIS SCUTATA

15 THYASIRA FLEXUOSA

16 ALVANIA CARINATA

17 SCOLOPLOS SP.

18 DULICHIA MONOCANTHA

19 TEREBELLIB A

20 OLIGOCHAETA

21 ARICIBEA SUECICA

22 AMPHARETE ARCTICA

23 EUBORELLA TRUNCATULA

24 LEPTOCHEIRUS PINGUIS

25 PERIPLOMA PAPYRATIUM

26 SABELLA PENICILLUS

27 IHPLOCIRRUS HIRSUTUS

28 NUCULA ANNULATA

29 APISTOBRANCHUS TULLBERGI

30 NEPHTYS INC ISA

31 MONOCULOIlES TESSELATUS

32 PARAONIS GRACILIS

33 RHODINE LOVENI

34 PHYLLOHOCE MUCOSA

35 ETEONE LONGA

36 OUENIA FUSIFORMIS

37 LUMBRINERIS TENUIS

38 YOLDIA LIMATULA

39 CEREBRATULUS LACTEUS

40 'CHAETOPTERUS SP.

41 TEREBELLIDES STROEMI

42 HARMATHOE IMBRICATA

43 PHERUSA AFFINIS

44 SCALIBREGMA INFLATUM

45 ARICIBEA OUABRILOBATA

46 SPIOPHANES BOMBYX

47 LAONICE CIRRATA

48 FHOLOE MINUTA

49 DIASTYLIS CORNUIFER

50 BIASTYLIS SCULPTA

51 BIASTYLIS OUABR ISPl'NOSA

52 LEPTOSTYLIS LONGIMANA

53 PETALOSARSIA BECLIVIS

54 HALIMEUON SP.

55 STENOPLEUSTES INERMIS

56 ANONYX LILJEBORGI

57 ARGISSA HAMATIPES

58 AMPELISCA MACROCEPHALA

59 CERASTOBERMA PINNULATUM

60 CYLICHNA GOULDI

61 CHIRIDOTA LAEVIS

62 ASTERIAS SP.

COUNT CUM COUNT

X

CUM 7.

165

165.

20. 17

20.17

87

252.

10.64

30.81

81

333.

9.90

40.71

78

411 .

9.54

50.24

49

460.

5.99

56.23

40

500.

4.89

61 .12

32

532.

3.91

65.04

25

557.

3.06

68.09

23

580.

2.81

70.90

21

601 .

2.57

73.47

20

621.

2.44

75.92

18

639.

2.20

78. 12

17

656.

2.08

80.20

16

672.

1.96

82.15

16

688.

1.96

84.11

15

703.

1.83

85.94

11

714.

1.34

87.29

7

721 .

0.86

88.14

6

727.

0.73

88.88

6

733.

0.73

89.61

5

738.

0.61

90.22

5

743.

0.61

90.83

5

748.

0.61

91 .44

5

753.

0.61

92.05

5

758.

0.61

92.66

4

762.

0.49

93.15

4

766.

0.49

93.64

4

770.

0.49

94.13

3

773.

0.37

94.50

3

776.

0.37

94.87

3

779.

0.37

95.23

2

781 .

0.24

95.48

i

783.

0.24

95.72

2

785.

0.24

95.97

2

787.

0.24

96.21

2

789.

0.24

96.45

2

791 .

0.24

96.70

2

793.

0.24

96.94

2

795.

0.24

97.19

796.

0. 12

97.31

797.

0.12

97.43

798.

0.12

97.55

799.

0. 12

97.68

800.

0.12

97.80

801.

0.12

97.92

802.

0.12

98.04

803.

0.12

98. 17

804.

0.12

98.29

805.

0.12

98.41

806.

0.12

98.53

807.

0.12

98.65

808.

0.12

98.78

809.

0. 12

98.90

810.

0.12

99.02

811.

0.12

99.14

812.

0. 12

99.27

813.

0.12

99.39

814.

0.12

99.51

815.

0.12

99.63

816.

0.12

99.75

817.

0.12

99.88

818.

0.12

100. 00

NUMBER OF SPECIES 62

NUMBER OF INI1I V IIJUALS BIB.

INUIVIDUALS PER M2 8180

127

CRUISE EX8001

STATION 11

RANK SPECIES NAME

1 PRIONOSFIO STEENSTRUPI

2 SPIO FILICORNIS

3 MEDIOMASTUS AMBISETA

4 PARAONIS GRACILIS

5 AMPELISCA VABORUM

6 ARCTICA ISLANUICA

7 NUCULA tiELPHINOHONTA

8 NINOE NIGRIPES

9 THYASIRA FLEXUOSA

10 STERNASPIS SCUTATA

11 THARYX SP.

12 EUPORELLA TRUNCATULA

13 PHOTIS MACROCOXA

14 SCOLOPLOS SP.

15 ALVANIA CARINATA

16 CEREBRATULUS LACTEUS

17 EDOTEA TRILOBA

18 AMPHARETE ARCTICA

19 CRENELLA DECUSSATA

20 OLIGOCHAETA

21 DIASTYLIS SCULPTA

22 SABELLA PENICILLUS

23 RHODINE LOVENI

24 HIPPOMEDON SERRATUS

25 STENOPLEUSTES INERMIS

26 PERIPLOMA PAPYRATIUM

27 NUCULA ANNULATA

28 PHOLOE MINUTA

29 ARICIDEA JEFFREYSII

30 NEPHTYS INCISA

31 EUHORELLA HISPIDA

32 HARPINIA PROPINGUA

33 ARRISSA HAMATIPES

34 METOPELLA ANGUSTA

35 MODIOLUS MODIOLUS

36 MYA ARENARIA

37 NUCULA TENUIS

38 PITAR MORRHUANA

39 DIPLOCIRRUS HIRSUTUS

40 LUMBRINERIS TENUIS

41 AGLAOPHAMUS NEOTENUS

42 AMPHARETE ACUTIFRONS

43 EUCLYMENE COLLARIS

44 ETEONE LONGA

45 LUMBRINFRIS FRAGILIS

46 BATHYMEDON SP.
4 7 HALIMEDOM SP.

48 MONOCULODES N.SP.

49 OXYUROSTYLIS SMITHI

50 PHOXOCEPHALUS HOLBOLLI

51 ORCHOMENELLA PINGUIS

C0UN1

CUM COUNT

%

CUM Z

505

505.

55 . 37

55.37

77.

582.

8.44

63.82

63

645.

6.91

70.72

27

672.

2.96

73.68

25

697.

2.74

76.43

20

717.

2.19

78.62

18

735.

1 .97

80.59

18

753.

1.97

82.57

17

770.

1.86

84.43

16

786.

1 .75

86.18

12

798.

1.32

87.50

12

810.

1.32

88.82

10

820.

1. 10

89.91

8

828.

0.88

90.79

7

835.

0.77

91.56

7

842.

0.77

92.32

6

848.

0.66

92.98

5

853.

0.55

93.53

4

857.

0.44

93.97

4

861 .

0.44

94.41

4

865.

0.44

94.85

3

868.

0.33

95.18

3

871.

0.33

95.50

3

874.

0.33

95.83

3

877.

0.33

96.16

2

879.

0.22

96.38

2

881.

0.22

96.60

2

883.

0.22

96.82

2

885.

0.22

97.04

2

887.

0.22

97.26

2

889.

0.22

97.48

2

891.

0.22

97.70

2

893.

0.22

97.92

2

895.

0.22

98.14

896.

0.11

98.25

897.

0. 11

98.35

898.

0. 11

98.46

899.

0.11

98.57

900.

0.11

98.68

901.

0.11

98.79

902.

0.11

98.90

903.

0.11

99.01

904.

0.11

99.12

905.

0.11

99.23

906.

0.11

99.34

907.

0.11

99.45

908.

0.11

99.56

909.

0. 11

99.67

910.

0. 11

99.78

911.

0.11

99.89

912.

0.11

100.00

NUMBER OF SPECIES 51

NUMBER OF INDIVIDUALS 912.
INDIVIDUALS PER M2 9120

128

CRUISE EXB001

STATION 12

RANK SPECIES NAME

1 PRIONOSPIO STEENSTRUPI

2 NUCULA PELPHINODONTA

3 MEIHOMASTUS AMBISETA

4 SPIO FILICORNIS

5 MAYERELLA LIMICOLA

6 STERNASPIS SCUTATA

7 NINOE NIGRIPES

8 EUDORELLA TRUNCATULA

9 ARICIDEA SUECICA

10 STENOPLEUSTES INERMIS

11 THYASIRA FLEXUOSA

12 SCOLOPLOS SP.

13 HAL I ME DON SP .

14 PARAONIS GRACILIS

15 PHOT IS MACROCOXA

16 ALBANIA CARINATA

17 DIASTYLIS SCULPTA

18 SABELLA PENICILLUS
1? DULICHIA MONOCANTHA

20 APISTOBRANCHUS rULLBERGl

21 ANEMONE A

22 MONOCULODES TESSELATUS

23 ANONYX LILJEBORGI

24 METOPELLA ANGUSTA

25 THARYX SP.

26 ETEONE LONGA

27 CEREBRATULUS LACTEUS

28 MYRIOCHELE HEERI

29 OLIGOCHAETA

30 AGLAOPHAMUS NEOTENUS

31 LUMBRINERIS TENUIS

32 LEPTOSTYLIS LONGIMANA

33 BATHYMEDON SP.

34 HARFINIA PROPINQUA

35 AMPELISCA AGASSIZI

36 ARGISSA HAMATIPES

37 DENTAL IUM ENTALE

38 NUCULA ANNULATA

39 ARCTICA ISLANUICA

40 PERIPLOMA PAPYRATIUM

41 MYA ARENARIA
4 2 PHOLOE MI NUT A

43 RHODINE LOVENI

44 EUCLYMENE COLLARIS

45 ARICIDEA QUADRILOBATA

46 ARICIDEA JEFFREYSII

47 PHYLLODOCE MUCOSA

48 PHERUSA AFFINIS

49 MOLPADIA OOLICTICA

50 NEMERTEA C

COUNT CUM COUNT

•/.

CUM 7.

351

351 .

55.36

5 j • 36

30

381 .

4.73

60.09

27

, 408.

4.26

64.35

23

431.

3.63

67.98

18

449.

2.84

70 . 82

17

466.

2.68

73.50

16

482.

-) ( ^o

76.03

15

497.

2.37

78.39

12

509.

1 .1)9

80.28

1 1

520.

1 . 74

82.02

10

530.

1 .58

83.60

9

539.

1 .42

85.02

8

e.47.

1 .26

86.28

8

555.

1 .26

87.54

6

561 .

0.95

88.49

6

567.

0.95

89.43

5

572.

0.79

90.22

5

577.

0.79

91 .01

4

581 .

0.63

91.64

4

585.

0.63

92.27

4

589.

0.63

92.90

3

592.

0.47

93.38

3

595.

0.47

93.85

3

598.

0.47

94.32

3

601.

0.47

94.79

3

604.

0.47

95 . 27

3

607.

0.47

95.74

->

609.

0.32

96.06

r>

61 1 .

0.32

96.37

•^

613.

0.32

96.69

2

615.

0.32

97.00

616.

0. 16

97.16

617.

0. 16

97.32

618.

0. 16

97.48

619.

0.16

97.63

620.

0. 16

97.79

621 .

0.16

97.95

622.

0. 16

98.11

623.

0. 16

98.26

624.

0.16

98.4 2

625.

0.16

98.58

626.

0. 16

98.74

627.

0. 16

98.90

628.

0. 16

99.05

629.

0. 16

99.21

630.

0.16

99.37

631 .

0. 16

99.53

632.

0.16

99.68

633.

0.16

99.84

634.

0. 16

100.00

NUMBER OF SPECIES 50

NUMBER OF INDIVIDUALS 634.
INDIVIDUALS PER M2 6340

CRUISE EX8001

STATION 13

RANK SPECIES NAME

1 PRIONOSPIO STEENSTRUPI

2 EUDORELLA TRUNCATULA

3 MEDIOMASTUS AMBISETA

4 SCOLOPLOS SP .

5 AGLAOPHAMUS NEOTENUS

6 ARICIHEA GUADRILOBATA

7 ARICIHEA JEFFREYSII

8 NUCULA DELPHINODONTA

9 ERYTHROPS ERYTHROPHTHALMA

10 CEREBRATULUS LACTEUS

11 NEPHTYS INCISA

12 ANEMONE A

13 OLIGOCHAETA

14 MAYERELLA LIMICOLA

15 EUDORELLA HISPIDA

16 CASCO BIGELOUI

17 LUMBRINERIS TENUIS

18 ETEONE LONGA

19 STAURONEREIS CAECUS

20 SPIO FILICORNIS

21 GEMMA GEMMA

22 ARGISSA HAMATIPES

23 MELITA N.SP.

24 ARCH I ANNELIDA

25 LUMBRINERIS FRAGILIS

26 BRADA VILLOSA

27 HARTMANIA MOOREI

28 APISTOBRANCHUS TULLBERGI

29 STERNASPIS SCUTATA

30 PARAOHIS GRACILIS

31 NEMERTEA H

32 MODIOLUS MODIOLUS

33 NUCULA ANNULATA

34 NASSARIUS TRIVITTATUS

35 DIASTYLIS SCULPTA

36 HALIMEDON SP.

37 PHOT IS MACROCOXA

38 HARP IN I A PROPINQUA

39 ORCHOMENELLA PINGUIS

40 AMPELISCA ABDITA

NUMBER OF SPECIES 40

NUMBER OF INDIVIDUALS 608.
INDIVIDUALS PER M2 6080

129

OUNT CUM COUNT

V.

CUM ■/.

267

267.

43.91

43.91

135

402.

22.20

66. 12

47

449.

7.73

73.85

43

492.

7.07

80.92

22

514.

3.62

84.54

14

528.

2.30

86.84

8

536.

1.32

88. 16

8

544.

1 .32

89.47

8

552.

1.32

90.79

5

557 .

0.82

91 .61

4

561 .

0.66

92.27

4

565.

0.66

92.93

3

568.

0.49

93.42

3

571.

0.49

93.91

3

574.

0.49

94.41

3

577.

0.49

94.90

2

579.

0.33

95.23

2

581.

0.33

95.56

2

583.

0.33

95.89

2

585.

0.33

96.22

2

587.

0.33

96.55

2

589.

0.33

96.87

2

591.

0.33

97.20

592.

0. 16

97.37

593.

0. 16

97.53

594.

0.16

97.70

595.

0.16

97.86

596.

0.16

98.03

597.

0.16

98.19

598.

0.16

98.35

599.

0.16

98.52

600.

0.16

98.68

601 .

0.16

98.85

602.

0.16

99.01

603.

0.16

99.18

604.

0.16

99.34

605.

0.16

99.51

606.

0.16

99.67

607.

0.16

99.84

608.

0.16

100.00

130

CRUISE EX8001

STATION I!

RANt^
1
2

3

4

5

6

7

8

9

10

11

12

13

14

15

16

17

IB

19

20

21

22

23

24

25

26

27

SPECIES NAME
PRIONOSPIO STEENSTRUPI
EUDORELLA TRUNCATULA
AGLAOPHAMUS NEOTENUS
MEIHOMASTUS AMBISE1A
SCOLOPLOS SP.
ARICIHEA SUECICA
[HASTYLIS SCULPTA
ANEMONE A
OLIGOCHAETA
CEREBRATULUS LACTEUS
EUPORELLA HISPIBA
BULICHIA MONOCANTHA
ARGISSA HAMATIPES
MELITA N.SP.
AMPELISCA ABDITA
PHASCOLOPSIS GOULIHI
NASSARIUS TRIVITTATUS
NEPHTYS INCISA
HAL I ME BON SP.
OUENIA FUSIFORMIS
LUMBRINERIS FRAGILIS
STERNASPIS SCUTATA
ARICIDEA OUAIiRILOBATA
THARYX SP.
LUMBRINERIS TENUIS
PHYLLODOCE MUCOSA
SPIO FILICORNIS

C0UN1

CUM COUNT

•/.

CUM X

112.

112.

22.67

22.67

99

211.

20.04

42.71

86

297.

17.41

60.12

74

371.

14.98

75.10

28

399.

5.67

80.77

25

424.

5.06

85.83

14

438.

2.83

88.66

10

448.

2.02

90.69

9

457.

1.82

92.51

6

463.

1.21

93.72

5

468.

1 .01

94.74

3

471.

0.61

95.34

3

474.

0.61

95.95

3

477.

0.61

96.56

2

479.

0.40

96.96

2

481.

0.40

97.37

2

483.

0.40

97.77

2

485.

0.40

98.18

486.

0.20

98.38

487.

0.20

98.58

488.

0.20

98.79

489.

0.20

98.99

490.

0.20

99.19

491.

0.20

99.39

492.

0.20

99.59

493.

0.20

99.80

. 494.

0.20

100.00

NUMBER OF SPECIES 27

NUMBER OF INUIUI IiUALS 494.
INDIVIDUALS PER M2 4940

CRUISE EX8001

STATION 16

RANK
1

3
4

6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24

SPECIES NAME
PRIONOSPIO STEENSTRUPI
AGLAOPHAMUS NEOTENUS
MEDIOMASTUS AMBISETA
SCOLOPLOS SP.
EUDORELLA TRUNCATULA
ANEMONE A
HIASTYLIS SCULPTA
LUMBRINERIS TENUIS
OLIGOCHAETA
NEOMYSIS AMERICANA
ERYTHROPS ERYTHROPHTHALMA
ARICIDEA SUECICA
OUENIA FUSIFORMIS
AMPELISCA ABDITA
STENOPLEUSTES INERMIS
NEPHTYS INCISA
LUMBRINERIS FRAGL 1LS
PHYLLOtlOCE MUCOSA
THARYX SP.
CYLICHNA ALBA
NASSARIUS TRIVITTATUS
CEREBRATULUS LACTEUS
OXYUROSTYLIS SMITHI
PHOT IS MACROCOXA

NUMBER OF SPECIES 24

NUMBER OF INDIVIDUALS 290.
INDIVIDUALS PER M2 2900

131

COUNT

CUM COUNT

V.

CUM X

100.

100.

34.48

34.48

62.

162.

21 .38

55.86

26.

188.

8.97

64.83

25.

213.

8.62

73.45

23.

236.

7.93

81.38

8

244.

2.76

84.14

8.

252.

2.76

86.90

6

258.

2.07

88.97

5

263.

1 .72

90.69

5

268.

1.72

92.41

4

272 .

1.38

93.79

3

275.

1 .03

94.83

2

277.

0.69

95.52

2

279.

0.69

96.21

2

281 .

0.69

96.90

282.

0.34

97.24

283.

0.34

97.59

284.

0.34

97.93

285.

0.34

98.28

286.

0.34

98.62

, 287.

0.34

98.97

288.

0.34

99.31

289.

0.34

99.65

290.

0.34

100.00

CRUISE EX8001

STATION 17

RANK SPECIES NAME

1 AGLAOPHAMUS NEOTENUS

2 ANEMONE A

3 MULINIA LATERALIS

4 NEPHTYS INCISA

5 PRIONOSPIO STEENSTRUPI

6 DULICHIA MONOCANTHA

7 NEOMYSIS AMERICANA

NUMBER OF SPECIES 7

NUMBER OF INDIVIDUALS 328.
INDIVIDUALS PER M2 3280

COUNT

CUM COUNT

V.

CUM X

276.

276.

84. 15

84. 15

40.

316.

12.20

96.34

7.

323.

2.13

98.48

2 ,

325.

0.61

99.09

1.

326.

0.30

99.39

1.

327.

0.30

99. 70

1.

328.

0.30

100.00

132

CRUISE EX8001

STATION 18

RANt

1

3

4

5

6

7

8

9

10

11

12

13

14

15

16

17

18

19

20

21

22

23

24

25

26

27

SPECIES NAME
OLIGOCHAETA
LUMBRINERIS TENUIS
ARICIDEA JEFFREYSII
LIMNORIA LIGNORUM
AMPHARETE ARCTICA
POLYHORA SOCIALIS
NINOE NIGRIPES
PHOT IS MACROCOXA
ETEONE LONGA
PRIONOSPIO STEENSTRUPI
PHOLOE MINUTA
GEMMA GEMMA
LUMBRINERIS FRAGILIS
TELLINA AGILIS
MEMBRANIPORIDAE
NEPHTYS INCISA
NEREIS VIRENS
YOLIHA LIMATULA
MYA ARENARIA
LYONSIA HYALINA
NASSARIUS TRIVITTATUS
COROPHIUM INSIDIOSUM
UN I COLA IRRORATA
SCOLOPLOS SP.
OPHELINA ACUMINATA
AGLAOPHAMUS NEOTENUS
CERIANTHUS BOREALIS

NUMBER OF SPECIES 27

NUMBER OF INDIVIDUALS 21 7.+
INDIVIDUALS PER M2 2170+

133

COUNT CUM COUNT

■/.

CUM 7.

43

43.

19.82

19.82

39

82.

17.97

37.79

37

119.

17.05

54.84

30

149.

13.82

68.66

14

163.

6.45

75.12

6

169.

2.76

77.88

6

175.

2.76

80.65

5

180.

2.30

82.95

5

185.

2.30

85.25

5

190.

2.30

87.56

4

194.

1 .84

89.40

,5

197.

1 .38

90.78

3

200.

1 .38

92.17

2

202.

0.92

93.09

2

204.

0.92

94.01

2

206.

0.92

94.93

2

208.

0.92

95.85

209.

0.46

96.31

210.

0.46

96.77

211.

0.46

97.23

212.

0.46

97.70

213.

0.46

98.16

214.

0.46

98.62

215.

0.46

99.08

216.

0.46

99.54

217.

0.46

100.00

CRUISE EX8001

STATION 19

RANK SPECIES NAME

1 LUMBRINERIS TENUIS

2 NINOE NIGRIPES

3 AGLAOPHAMIIS NEOTENUS

4 NUCULA DELPHINODONTA

5 PRIONOSPIO STEENSTRUPI

6 PHYLLODOCE MUCOSA

7 EUDORELLA TRUNCATULA

8 ORCHOMENELLA PINGUIS

9 ARICIDEA JEFFRErSII

10 AMPHARETE ARCTICA

11 OLIGOCHAETA

12 TELLINA AGILIS

13 NEMERTEA C

14 PHOLOE MINUTA

15 YOLDIA LIMATULA

16 ETEONE LONGA

17 PHERUSA AFFINIS
IB NEPHTYS INCISA

19 METERYTHROPS ROBUSTA

20 MEDIOMASTUS AMBISETA

21 LUMBRINERIS FRAGILIS

22 POTAMILLA NEGLECTA

23 NEREIS SP.

24 GEMMA GEMMA

25 NUCULA ANNULATA

26 MODIOLUS MODIOLUS

27 ANEMONE A

28 NEMERTEA E

29 CERIANTHUS BOREALIS

30 DIASTYLIS SCULPTA

31 AMPELISCA ABDITA

32 LEPTOCHEIRUS PINGUIS

NUMBER OF SPECIES 32

NUMBER OF INDIVIDUALS 260.
INDIVIDUALS PER M2 -> 60

134

COUNT CUM COUNT

7.

CUM 7.

73

73.

28.08

28.08

31

104.

11 .92

40.00

30

134.

11 .54

51 .54

19

153.

7.31

58.85

15

168.

5.77

64.62

13

181.

5.00

69.62

11

192.

4.23

73.85

10

202.

3.85

77.69

9

211.

3.46

81.15

8

219.

3.08

84.23

6

225.

2.31

86.54

4

229 .

1 .54

88.08

A

233.

1 .54

89.62

3

236.

1.15

90.77

3

239.

1.15

91 .92

O

241.

0.77

92.69

2

243.

0.77

93.46

2

245.

0.77

94.23

2

247.

0.77

95.00

248.

0.38

95.38

249.

0.38

95.77

250.

0.38

96. 15

251.

0.38

96.54

2S2.

0.38

96.92

253.

0.38

97.31

254.

0.38

97.69

255.

0.38

98.08

256.

0.38

98.46

257.

0.38

98.85

258.

0.38

99.23

259.

0.38

99.62

260.

0.38

100.00

CRUISE EX8001

STATION 20

RANK SPECIES NAME

1 NUCULA DELPHINODONTA

2 PRIONOSPIO STEENSTRUPI

3 EUDORELLA TRUNCATULA

4 MEDIOMASTUS AMBISE1A

5 LUMBRINERIS TENUIS

6 ANEMONE A

7 AGLAOPHAMUS NEOTENUS

8 NINOE NIGRIPES

9 THARYX SP.

10 IHASTYLIS SCULPTA

11 ARGISSA HAMATIPES

12 OLIGOCHAETA

13 OUENIIDAE

11 CEREBRATULUS LACTEUS

15 PHYLLODOCE MUCOSA

16 GEMMA GEMMA

17 HYDRO? I A SP.

18 NASSARIUS TRIVITTATUS

19 ORCHOMENELLA PINGUIS

20 LEPTOCHEIRUS PINGUIS

21 ARICIDEA JEFFREYSII

22 YOLDIA LIMATULA

23 NUCULA ANNULATA

24 CERASTODERMA PINNULATUM

25 TELLINA AGILIS

26 NEOMYSIS AMERICANA

27 NEMERTEA C

NUMBER OF SPECIES 27

NUMBER OF INOI VI HUALS 298.
INDIVIDUALS PER M2 2980

135

COUNT

CUM COUNT

■/.

CUM 7.

113.

113.

37.92

37.92

68.

181 .

22.82

60.74

18.

199.

6.04

66.78

15.

214.

5.03

71.81

15

229.

5.03

76.85

11

240.

3.69

80.54

7

247.

2.35

82.89

6

253.

2.01

84.90

5

258.

1 .68

86.58

4

262.

1 .34

87 . 92

4

266.

1.34

89.26

4

270.

1.34

90.60

4

274.

1.34

91.95

3

277,

1 .01

92.95

3

280.

1.01

93.96

2

282.

0.67

94.63

2

284.

0.67

95 . 30

2

286.

0.67

95.97

2

288.

0.67

96.64

2

290.

0.67

97.32

2

292.

0.67

97.99

293.

0.34

98.32

294.

0.34

98.66

295.

0.34

98.99

296.

0.34

99.33

297.

0.34

99.66

298.

0.34

100.00

136

CRUISE EX8001

STATION 21

GRAB 1

RANN SPECIES NAME

1 NUCULA DELPHINODONTA

2 PRIONOSPIO STEENSTRUPI

3 LUMBRINERIS TENUIS

4 ARICIDEA JEFFREYSII

5 MEDIOMASTUS AMBISETA

6 SCOLOPLOS SP.

7 EUDORELLA TRUNCATULA

8 PHOXOCEPHALUS HOLBOLLI

9 AGLAOPHAMUS NEOTENUS

10 DIASTYLIS SCULPTA

11 DRCHOMENELLA PINGUIS

12 NINOE NIGRIPES

13 PHOTIS MACROCOXA

14 OLIGOCHAETA

15 PHYLLOHOCE MUCOSA

16 PITAR MORRHUANA

17 BA1HYMFD0N SP .

18 DULICHIA MONOCANTHA

19 CEREBRATULUS LACTEUS

20 THARYX SP.

21 ETEONE LONGA

22 PERIPLOMA PAPYRATIUM

23 CERIANIHUS BOREALIS

24 ARGISSA HAMATIPES

25 NEMERTEA C

26 AMPHARETE ACUTIFRONS

27 POTAMILLA NEGLECTA

28 MY A ARENARIA

29 NUCULA ANNULATA

30 CHIRIDOTA LAEVIS

31 STENOPLEUSTES INERMIS

32 CASCO BIGELOUI

33 SABELLA PENICILLUS

34 OUENIA FUSIFORMIS

35 PHERUSA AFFINIS

36 NEPHTYS INCISA

37 CAPITELLA CAPITATA

38 PARAONIS GRACILIS

39 STAURONEREIS CAECUS

40 MODIOLUS MODIOLUS

41 CRENELLA DECUSSATA

COUNT CUM COUNT

•/.

CUM 7.

'.40

540.

35.79

35.79

335

875.

22.20

5 7.9?

182

1057.

12.06

70.05

92

1147.

6.10

76.14

53

1202.

3.51

79.66

49

1251.

3.25

82.90

38

1289.

2.52

85.4 2

36

1325.

2.39

87.81

31

1356.

2.05

89.86

29

1385.

1 .92

91 .78

18

1403.

1 . 19

92.98

17

1420.

1 . 13

94 . 10

14

1434.

0.93

95.03

14

1448.

0.93

95.96

5

1453.

0.33

96.29

5

1458.

0.33

96.62

4

1462.

0.27

96.89

4

1466.

0.27

97. 15

4

1470.

0.27

97.42

4

1474.

0.27

97.68

4

1478.

0.27

97.95

4

1482.

0.27

9FJ.21

3

1485.

0.20

98.41

2

1487.

0.13

98.54

2

1489.

0. 13

98.67

2

1491 .

0.13

9H.81

2

1493.

0.13

98.94

2

1495.

0. 13

99.07

2

1497.

0. 13

99.20

1498.

0.07

99.27

1499.

0.07

99.34

1500.

0.07

99.40

1501 .

0.07

99.47

1502.

0.07

99.54

1503.

0.07

99.60

1504.

0.07

99.67

1505.

0.07

99.73

1506.

0.07

99.80

1507.

0.07

99.87

1508.

0.07

99.93

1509.

0.07

100.00

NUMBER OF SPECIES

41

NUMBER OF INDIVIDUALS 1509.
INDIVIDUALS PER M2 15090

137

CRUISE EX8001

STATION 22

RANK SPECIES NAME

1 PRIONOSPIO STEENSTRUPI

2 MEDIOMASTUS AMBISETA

3 LUMBRINERIS TENUIS

4 ARICIDEA JEFFREYSII

5 AMPHARETE ARCTICA

6 NUCULA DELPHI NODONTA

7 EUDORELLA TRUNCATULA

8 NINOE NIGRIPES

9 THARYX SP.

10 CRENELLA HECUSSATA

11 PHOXOCEPHALUS HOLBOLLI

12 NASSARIUS TRIVITTATUS

13 ARCHIANNELIUA

14 AMPELISCA VADORUM

15 COROPHIUM CRASSICORNE

16 SCOLOPLOS SP.

17 DIASTYLIS SCULPTA

18 OLIGOCHAETA

19 POLYDORA SP .

20 UNCIOLA IRRORATA

21 ETEONE LONGA

22 PHOLOE MINUTA

23 HARP INI A PROPINQUA

24 LEPTOCHEIRUS PINGUIS

25 CERASTODERMA PINNULATUM

26 SYLLIS GRACILIS

27 AUTOLYTUS SP .

28 PI TAR MORRHUANA

29 POLYDORA SOCIAL IS

30 OPHELINA ACUMINATA

31 PHERUSA AFFINIS

32 NEMERTEA F

33 NEMERTEA C

34 ORCHOhENELLA PINGUIS

35 EUCLYMENE COLLARIS

36 SCOLOPLOS ROBUSTUS

37 CHIRIDOTA LAEVIS

38 NEMERTEA G

39 CEREBRATULUS LACTEUS

40 PROTOMEHEIA FASCIATA

41 GEMMA GEMMA

42 PERIPLOMA PAPYRATIUM

43 TELLINA AGILIS

44 HYDROBIA SP.

45 NEREIS PELAGICA

46 AMPHIPHOLIS SQUAMATA

47 CERIANTHUS BOREALIS

48 CANCER BOREALIS

49 PAGURUS PUBESCENS

50 STENOPLEUSTES GRACILIS

51 STENOPLEUSTES INERMIS

52 EDOTEA TRILOBA

53 DOTO CORONATA

54 COCCULINA SP .

55 ASTARTE UNDATA

56 PANDORA GOULDIANA

57 LYONSIA HYALINA

58 EXOGONE HEBES

59 POLYDORA QUADRILOBATA

60 LUMBRINERIS FRAGILIS

61 SPIOPHANES BOMBYX

62 MEMBRANIPORIDAE

63 SERTULARIA PUMILA

COUNT

CUM LillINT

V.

CUM 7.

269.

269.

28.50

28.50

120.

389.

12.71

41.21

71.

460.

7.52

48.73

59.

519.

6.25

54.98

54.

573.

5.72

60.70

48

621.

5.08

65.78

29

650.

3.07

68.86

28

678.

2.97

71.82

26

704.

2.75

74.58

23

727.

2.44

77.01

19

746.

2.01

79.03

17

763.

1 .80

80.83

15

778.

1 .59

82.42

12

790.

1 .27

83.69

10

800.

1.06

84.75

10.

810.

1 .06

85.80

9

819.

0.95

86.76

8

827.

0.85

87.61

8

835.

0.85

88.45

7

842.

0.74

89.19

7

849.

0.74

89.94

6

855.

0.64

90.57

5

860.

0.53

91 . 10

5

865.

0.53

91.63

5

870.

0.53

92. 16

5

875.

0.53

92.69

4

879.

0.42

93.11

4

883.

0.42

93.54

4

887.

0.42

93.96

4

891 .

0.42

94.39

4

895.

0.42

94.81

3

898.

0.32

95.13

3

901.

0.32

95.44

3

904.

0.32

95.76

3

907.

0.32

96.08

3

910.

0.32

96.40

2

912.

0.21

96.61

2

914.

0.21

96.82

2

916.

0.21

97.03

2

918.

0.21

97.25

2

920.

0.21

97.46

2

922 .

0.21

97.67

2

924.

0.21

97.88

2

926.

0.21

98.09

2

928.

0.21

98.30

929.

0. 11

98.41

930.

0.11

98.52

931 .

0.11

98.62

932.

0.11

98.73

933.

0.11

98.83

934.

0.11

98.94

935.

0.11

99.05

936.

0. 11

99.15

937.

0.11

99.26

938.

0.11

99.36

939.

0.11

99.47

940.

0. 11

99.58

941.

0. 11

99.68

942.

0.11

99.79

943.

0. 11

99.89

944.

0. 11

100.00

+

+

NUMBER OF SPECIES

63

NUMBER OF INDIVIDUALS 944. t
INDIVIDUALS PER M2 9440+

138

CRUISE EX8001

STATION 23

GRAB 1

RANK SPECIES NAME

1 PRIONOSPIO STEENSTRUPI

2 AGLAOPHAMUS NEOTENUS

3 LUMBRINERIS TENUIS

4 NUCULA DELPHINODONTA

5 EUIlORELLA TRUNCATULA

6 ARICIDEA JEFFREYSII

7 MEDIOMASTUS AMBISETA

8 ORCHOMENELLA PINGUIS

9 IiIASTYLIS SCULPTA

10 NINOE NIGRIPES

11 OLIGOCHAETA

12 AMPELISCA ABDITA

13 AMPHARETE ARCTICA

14 ARGISSA HAMATIPES

15 STAURONEREIS CAECUS

16 CASCO BIGELOUI

17 NEPHTYS INCISA

18 HARPINIA PROPINOUA

19 PHOXOCEPHALUS HOLBOLLI

20 CERIANTHUS BOREALIS

21 ETEONE LONGA

22 SCOLOPLOS SP.

23 LEPTOCHEIRUS PINGUIS

24 BATHYMEDON SP.

25 AMPHARETE ACUTIFRONS

26 SABELLA PENICILLUS

27 PHYLLODOCE MUCOSA

28 MICROPHTHALMUS ABERRANS

29 PITAR MORRHUANA

30 CRENELLA DECUSSATA

31 NEOMYSIS AMERICANA

32 METERYTHROPS ROBUSTA

33 COROPHIUM CRASSICORNE

34 MONOCULODES N.SP.

35 NEMERTEA C

36 SYLLIS GRACILIS

37 CLYMENELLA TORQUATA

38 PHERUSA AFFINIS

39 PARAONIS GRACILIS

40 PHOLOE MINUTA

41 YOLDIA LIMATULA

42 GEMMA GEMMA

COUNT CUM COUNT

X

CUM 7.

638

638.

44.55

44.55

180

818.

12.57

57.12

173

991.

12.08

69.20

156

1147.

10.89

80. 10

65

1212.

4.54

84.64

35

1247.

2.44

87.08

29

1276.

2.03

89.11

20

1296.

1 .40

90.50

17

1313.

1 . 19

91 .69

16

1329.

1.12

92.81

16

1345.

1 .12

93.92

11

1356.

0.77

94.69

9

1365.

0.63

95.32

7

1372.

0.49

95.81

6

1378.

0.42

96.23

5

131:3.

0.35

96.58

5

1388.

0.35

96.93

4

1392.

0.28

97.21

3

1395.

0.21

97.42

3

1398.

0.21

97.63

3

1401 .

0.21

97.83

3

1404.

0.21

98.04

2

1406.

0. 14

98. 18

2

1408.

0. 14

98.32

2

1410.

0. 14

98.46

2

1412.

0.14

98.60

2

1414.

0.14

98.74

2

1416.

0.14

98.88

2

1418.

0.14

99.02

2

1420.

0.14

99. 16

1421 .

0.07

99.23

1422.

0.07

99.30

1423.

0.07

99.37

1424.

0.07

99.44

1425.

0.07

99.51

1426.

0.07

99.58

1427.

0.07

99.65

1428.

0.07

99.72

1429.

0.07

99.79

1430.

0.07

99.86

1431.

0.07

99.93

1432.

0.07

100.00

NUMBER OF SPECIES

42

NUMBER OF INDIVIDUALS 1432.
INDIVIDUALS PER M2 14320

139

CRUISE EX8001

STATION 24

RANK

SFEC1ES NAME

1

AGLAOPHAMUS NEOTENUS

2

PRIONOSPIO STEENSTRUPI

3

ARICI&EA JEEFREYSII

4

MEIHOMASTUS AMBISETA

5

CASCO BIGELOUI

6

OLIGOCHAETA

7

EUUORELLA TRUNCATULA

8

ORCHOMENELLA PINGUIS

9

NINOE NIGRIPES

10

LEPTOCHEIRUS PINGUIS

11

ETEONE LONGA

12

AMPELISCA ABDITA

13

PHOXOCEFHALUS HOLBOLLI

14

SCOLOPLOS SP.

15

LUMBRINERIS TENUIS

16

NEPHTYS INC ISA

17

THARYX SF'.

18

NEMERTEA H

19

[IULICHIA MONOCANTHA

20

UNCI OLA IRRORATA

21

NEREIS VIRENS

22

PHYLLODOCE MUCOSA

23

POLYHORA LIGNI

24

NEMERTEA C

25

[iIASTYLIS SCULPTA

26

PHOT IS MACROCOXA

27

NEOMYSIS AMERICANA

NUMBER OF SPECIES 27

NUMBER OF INDI VI HUALS 873.
INUIV1UUALS PER M2 8730

COUNT CUM COUNT

•/.

CUM 7.

T>27

227.

26.00

26.00

206

433.

23.60

49.60

196

629.

22.45

72.05

77

706.

8.82

80.87

40

746.

4.58

85. 45

31

777.

3.55

89.00

18

795.

2.06

91.07

16

811.

1 .83

92.90

13

824.

1.49

94.39

10

834.

1.15

95.53

6

840.

0.69

96.22

6

846.

0.69

96.91

4

850.

0.46

97.37

3

853.

0.34

97.71

3

856.

0.34

98.05

2

858.

0.23

98.28

2

860.

0.23

98.51

2

862.

0.23

98.74

2

864.

0.23

98.97

2

866.

0.23

99.20

867.

0.11

99.31

868.

0. 11

99.43

869.

0.11

99.54

870.

0.11

99.66

871 .

0. 11

99.77

872.

0.11

99.89

873.

0.11

100.00

CRUISE EX8001

STATION 21

RANK SPECIES NAME

1 AGLAOPHAMUS NEOTENUS

2 ANEMONE A

3 CASCO BIGELOWI

4 PRIONOSPIO STEENSTRUPI

5 DIASTYLIS SCULF'TA

6 EUHORELLA TRUNCATULA

7 MEDIOMASTUS AMBISETA

8 ARICIDEA JEFFREYSII

9 NEOMYSIS AMERICANA

10 METERYTHROPS ROBUSTA

11 OLIGOCHAETA

12 POTAMILLA NEGLECTA

13 ORCHOMENELLA PINGUIS

14 AMPELISCA ABDITA

15 MELITA N.SP.

16 LUMBRINERIS TENUIS

17 NINOE NIGRIPES

18 ERYTHROPS ERYTHROPHTHALMA

19 ARGISSA HAMATIPES

20 YOLDIA LIMATULA

NUMBER OF SPECIES

20

NUMBER OF INDIVI DUALS 426 .
INDIVIDUALS PER M2 ■'260

140

COUNT Clin COUNT

V.

CUM 7.

219

219.

51 .41

51 .41

50

269.

11 .74

63. 15

47

316.

1 1 .03

74. 18

37

353.

8.69

82.86

19

372.

4.46

87.32

14

386.

3.29

90.61

8

394.

1 .88

92.49

7

401.

1.64

94. 13

7

408.

1 .64

95.77

3

411 .

0.70

96.48

2

413.

0.47

96.95

2

415.

0.47

97.42

2

417.

0.47

97.89

2

419.

0.47

98.36

2

421.

0.47

98.83

422.

0.23

99.06

423.

0.23

99.30

424.

0.23

99.53

425.

0.23

99.77

426.

0.23

100.00

CRUISE EX8001

STATION 26

RANK

1

2

3

4

5

6

7

8

9

10

11

12

13

14

15

16

1 7

18

SPECIES NAME
AGLAORHAMUS NEOTENUS
ANEMONE A

PRIONOSPIO STEENSTRUPI
ARICIHEA JEFFREYSII
MEBIOMASTUS AMBISETA
EUHORELLA TRUNCATULA
NEOMYSIS AMERICANA
IiULICHIA MONOCANTHA
ERYTHROPS ERYTHROPHTHAl Mi"
LITTORINA OBTUSATA
OLIGOCHAETA
NEPHTYS INC ISA
METERYTHROPS ROBUSTA
COROPHIUM CRASSICORNE
HARP INI A PROPINQUA
MELITA N.SP.
PHOT IS MACROCOXA
YOLDIA LIMATULA

NUMBER OF SPECIES

18

NUMBER OF INDIV I tlUALS 150.

INDIVIHUALS PER M2 1500

141

COUNl

CUM COUNT

•/.

CUM X

98

98.

65.33

65.33

12

110.

8.00

73.33

8

118.

5.33

78.67

7

125.

4.67

83.33

5

130.

3.33

B6.67

3

133.

2.00

88.67

3

136.

2.00

90.67

2

138.

1.33

92.00

2

140.

1.33

93.33

2

142.

1 .33

94.67

143.

0.67

95.33

144.

0.67

96.00

145.

0.67

96.67

146.

0.67

97.33

147.

0.67

98.00

148.

0.67

98.67

149.

0.67

99.33

150.

0.67

100.00

CRUISE EX8001

STATION 27

RANK

1

5

3
4

5
6
7
8
9

10
11
12
13
14
15
16
17
18
1?
20
21
22
23
24
25
26

SPECIES NAME
PRIONOSPIO STEENSTRUPI
EUDORELLA TRUNCATULA
ARICIDEA JEFFREYSII
NINOE N I GRIPES
NUCULA DELPHINODONTA
SCOLOPLOS SP.
AGLAOPHAMUS NEOTENUS
MEDIOMASTUS AMBISETA
OLIGOCHAETA
ARICIDEA SUECICA
ERYTHROPS ERYTHROPHTHALHA
DIASTYLIS SCULPTA
ARGISSA HAMATIPES
OWENIA FUSIFORMIS
CEREBRATULUS LACTEUS
NEPHTYS INCISA
MONOCULODES TUBERCULATUS
METERYTHROPS ROBUSTA
MYSIS STENOLEPIS
MODIOLUS MODIOLUS
YOLDIA LIMATULA
MOLPADIA OOLICTICA
PARAONIS GRACILIS
THARYX SP.
LUMBRINERIS TENUIS
SPIO FILICORNIS

NUMBER OF SPECIES 26

NUMBER OF INDIVIDUALS 227.
INDIVIDUALS PER M2 2270

142

COUNT CUM COUNT

7.

CUM >.

112

112.

49.34

49.34

18

130.

7.93

57.27

16

146.

7.05

64.32

14

160.

6.17

70. 4E

10

170.

4.41

74.89

7

177.

3.08

77.97

7

184.

3.08

81 .06

6

190.

2.64

83.70

5

195.

2.20

85.90

5

200.

2.20

88.11

4

204.

1.76

89.87

3

207.

1.32

91.19

3

210.

1.32

92.51

3

213.

1.32

93.83

2

215.

0.88

94.71

2

217.

0.88

95.59

218.

0.44

96.04

219.

0.44

96.48

220.

0.44

96.92

221.

0.44

97.36

222.

0.44

97.80

223.

0.44

98.24

224.

0.44

98.68

225.

0.44

99. 12

226.

0.44

99.56

227.

0.44

100.00

143

CRUISE EX8001

STATION 28

GRAB 1

RANK SPECIES NAME

1 PRI0NOSPI0 STEENSTRUPI

2 LEFTOCHEIRUS PINGUIS

3 NUCULA PELPHINODONTA

4 EUDORELLA TRUNCATULA

5 LUMBRINERIS TENUIS

6 DIASTYLIS SCULPTA

7 NINOE NIGRIPES

8 ORCHOMENELLA PINGUIS

9 ARGISSA HAMATIPES

10 MEDIOMASTUS AMBISETA

11 AGLAOPHAMUS NEOTENUS

12 BATHYMEDON SP .

13 ARICIDEA JEFFREYSII

14 PHYLLODOCE MUCOSA

15 NEMERTEA D

16 PHOXOCEPHALUS HOLBOLLI

17 PHOT IS MACROCOXA

18 ARICIDEA SUECICA
1? PARAONIS GRACILIS

20 OPHELINA ACUMINATA

21 NEPHTYS INCISA

22 CEREBRATULUS LACTEUS

23 NEMERTEA C

24 HARPINIA PROPINOUA

25 STENOPLEUSTES INERMIS

26 DULICHIA MONOCANTHA

27 SCOLOPLOS SP.

28 SPIO FILICORNIS

29 NEMERTEA H

30 MAYERELLA LIMICOLA

31 MELITA N.SP.

32 PHOLOE MINUTA

33 OLIGOCHAETA

34 PLATYHELMINTHES

35 HALIMEDON SP.

36 METOPELLA ANGUSTA

37 UNCIOLA IRRORATA

38 STAURONEREIS CAECUS

39 SPIO SETOSA

40 TEREBELLID B

41 HARTMANIA MOORE I

42 THARYX SP .

43 GONIADA MACULATA

44 PITAR MORRHUANA

45 NUCULA ANNULATA

COUNT

CUM COUNT

7.

CUM V.

1088.

1088.

63.74

63.74

98.

1186.

5.74

69.48

83.

1269.

4.86

74.34

60.

1329.

3.51

77.86

60.

1389.

3.51

81 .37

52.

1441 .

3.05

84.42

47.

1488.

2.75

87.17

45.

1533.

2.64

89.81

22.

1 555 •

1.29

91 .10

15.

1570.

0.88

91 .97

15.

1585.

0.88

92.85

13.

1598.

0.76

93.61

12.

1610.

0.70

94.32

9.

1619.

0.53

94.84

8.

1627.

0.47

95.31

8.

1635.

0.47

95.78

7.

1642.

0.41

96.19

6.

1648.

0.35

96.54

5 .

1653.

0.29

96.84

5 *

1658.

0.29

97. 13

5.

1663.

0.29

97.42

4.

1667.

0.23

97.66

3.

1670.

0.18

97.83

3.

1673.

0.18

98.01

3.

1676.

0. 18

98.18

3.

167V.

0. 18

98.36

3.

1682.

0. 18

98.54

3.

1685.

0.18

98.71

2.

1687.

0.12

98.83

2.

1689.

0.12

98.95

2.

1691 .

0.12

99.06

2.

1693.

0.12

99.18

2.

1695.

0.12

99.30

1 .

1696.

0.06

99.36

1 .

1697.

0.06

99.41

1 .

1698.

0.06

99.47

1 .

1699.

0.06

99.53

1 .

1700.

0.06

99.59

1 .

1701.

0.06

99.65

1 .

1702.

0.06

99.71

1 .

1703.

0.06

99.77

1 .

1704.

0.06

99.82

1 .

1705.

0.06

99.88

1 .

1706.

0.06

99.94

1 .

1707.

0.06

100.00

NUMBER OF SPECIES 45

NUMBER OF INDI VIDUALS 1707.
INDIVIDUALS PER M2 17070

CRUISE EX8001

STATION 29

RANK SPEC II S NAME

1 NUCULA UELF'HINODONTA

2 PRIONOSPIO STEENSTRURI

3 EUHORELLA TRUNCATULA

4 NINOE NIGRIF'ES

5 DIASTYLIS SCULPTA

6 LUMBRINERIS TENUIS

7 CEREBRATULUS LACTEUS

8 AMPHARETE ACUTIFRONS

9 SCOLOPLOS SR.

10 NEPHTYS INCISA

11 AGLAOPHAMUS NEOTENUS

12 ARICIDEA SUECICA

13 SFIO FILICORNIS

14 MEPIOMASTUS AMBISETA

15 PARAONIS GRACILIS

16 EUHORELLA HISPIIiA

17 ARGISSA HAMATIPES

18 MELITA N.SP.
1? CASCO BIGELOUI

20 ERYTHROPS ERYTHROPHTHALMA

21 YOLDIA LIMATULA

22 THYASIRA FLEXUOSA

23 NUCULA ANNULATA

24 NASSARIUS TRIVITTATUS

25 MOLPAUIA OOLICTICA

NUMBER OF SPECIES 25

NUMBER OF INDI VI HUALS 358.
INDIVIUUALS PER M2 3580

144

COUN

CUM COUNT

7.

CUM 7.

1 10

110.

30.- '3

30.73

97

207.

27.09

57.8?

68

275.

18.99

76.82

33

308.

9.22

86.03

11

319.

3.07

89. 11

8

327.

2 . ^3

91 .34

4

331 .

1.12

92.46

3

334.

0.FJ4

93.30

3

337.

0.84

94.13

3

340.

0.84

94.97

2

342.

0.56

95.53

2

344.

0.56

96.09

?

346.

0.56

96.65

347.

0.28

96.93

348.

0.28

97.21

349.

0.28

97.49

350.

0.28

97.77

351 .

0.28

98.04

352.

0.28

98.32

353.

0.28

98.60

354.

0.28

98.FJ8

355.

0.28

99.16

356.

0.28

99.44

357.

0.28

99.72

358.

0.28

100.00

145

CRUISE EX8001

STATION 30

RANK

SPECIES NAME

1

EUUORELLA TRUNCATULA

1

PR10N0SPI0 STEENSTRUPI

3

NINOE NIGRIPES

4

ERYTHROPS ERYTHROPHTHALMA

5

SPIO FILICORNIS

6

CEREBRATULUS LACTEUS

7

MELITA N.SP.

8

SCOLOPLOS SP.

9

DIASTYLIS SCULPTA

10

MEIHOMASTUS AMBISETA

11

NEPHTYS INCISA

12

GONIADA MACULATA

13

ARICIDEA SUECICA

14

ARGISSA HAMATIPES

NUMBER

OF SPECIES 14

NUMBER

OF INDIVIDUALS 108.

INDIVIDUALS PER M2 1080

COUNT CUM COUNT

CUM 7.

43.

43.

39.81

39.81

29.

72.

26.85

66.67

6.

78.

5.56

72.22

5.

83.

4.6*

76.85

4.

87.

3.70

80.56

4.

91.

3.70

84.26

4.

95.

3.70

87 . 96

3.

98.

2.78

90.74

3.

101.

2.78

93.52

2.

103.

1 .85

95.37

2.

105.

1.85

97.22

1 .

106.

0.93

98.15

1.

107.

0.93

99.07

1.

108.

0.93

100.00

CRUISE EX8001

STATION 31

RANK SPECIES NAME

1 PRIONOSPIO STEENSTRUPI

2 NINOE NIGRIPES

3 NEOMYSIS AMERICANA

4 NEPHTYS INCISA

5 SCOLOPLOS SP.

6 MEDIOMASTUS AMBISETA

7 CEREBRATULUS LACTEUS

8 ARICItiEA SUECICA

9 AGLAOPHAMUS NEOTENUS

10 EUDORELLA TRUNCATULA

11 DIASTYLIS SCULPTA

12 OLIGOCHAETA

13 ARGISSA HAMATIPES

14 DEXAMINE THEA

NUMBER OF SPECIES 14

NUMBER OF INDIVIDUALS 58.
INDIVIDUALS PER M2 580

146

COUNT CUM COUNT

X

CUM V.

23.

23.

39.66

39.66

6.

29.

10. 34

50.00

5.

34.

8.62

58.62

4.

38.

6.90

65.52

3.

41.

5.17

70.69

3.

44.

5.17

75.86

3.

47.

5.17

81 .03

2.

49.

3.45

84.48

2.

51.

3.45

87.93

2.

53.

3.45

91.38

2.

55.

3.45

94.83

1 .

54.

1.72

96.55

1.

57.

1.72

98.28

1.

58.

1.72

100.00

147

CRUISE EX8001

STATION 32

RANK
1

SPECIES NAME
AGLAOPHAMUS NEOTENUS
PRIONOSPIO STEENSTRUP1
DULICHIA MONOCANTHA
CASCO BIGELOUI
BIVALVIA

COUNT

CUM

COUNT

V.

CUM V.

100.

100.

94.34

94.34

3.

103.

2.83

97.17

1.

104.

0.94

98.11

1.

105.

0.94

99.06

1.

104.

0.94

100.00

NUMBER OF SPECIES 5

NUMBER OF INDIVIDUALS 106.
INDIVIDUALS PER M2 1060

CRUISE EX8001

STATION 33

RANK SPECIES NAME

1 PRIONOSPIO STEENSTRUPI

2 EUDORELLA TRUNCATULA

3 AGLAOPHAMUS NEOTENUS

4 NUCULA IiELPHINOnONTA

5 HIASTYLIS SCULPTA

6 ARICIDEA JEFFREYSII

7 MEDIOMASTUS AMBISETA

8 LUMBRINERIS TENUIS

9 OLIGOCHAETA

10 NINOE NIGRIPES

11 SCOLOPLOS SP.

12 ERYTHROPS ERYTHROPHTHALMA

13 YOLIHA LIMATULA
11 ARGISSA HAMATIPES

15 NEMERTEA H

16 CEREBRATULUS LACTEUS

17 THARYX SP.

18 MYRIOCHELE HEER1
1? NEPHTYS INCISA

20 ORCHOMENELLA PINGUIS

21 NEMERTEA D

22 MICROPHTHALMUS ABERRANS

23 ETEONE FLAVA

24 AMPHARETE ARCTICA

25 MALDANE SARSI

26 HARTMANIA MOORE I

27 ETEONE HETEROPOIiA

28 CASSIDINIHEA LUNIFRONS

29 DULICHIA MONOCANTHA

30 HALIMEDON SP .

31 GEMMA GEMMA

32 HYDROBIA SP .

NUMBER OF SPECIES 32

NUMBER OF INDIVIDUALS 925.

INDIVIDUALS PER M2

148

COUNT CUM COUNT

7.

CUM ■/.

434

434.

46.92

46.92

94

528.

10. 16

57.08

71

599.

7.68

64.76

61

660.

6.59

71.35

55

715.

*i . 95

77.30

52

767.

5.62

82.92

37

804.

4.00

86.92

24

828.

2.59

89.51

17

845.

1 .84

91 .35

17

as2.

1 .84

93.19

16

878.

1.73

94.92

12

890.

1 .30

9^.22

6

896.

0.65

96.86

4

900.

0.43

97.30

3

903.

0.32

97.62

2

905.

0.22

97.84

2

907.

0.22

98.05

2

909.

0.22

98.27

2

911 .

0.22

98.49

2

91 *

0.22

98.70

914,

0.11

98.81

915.

0.11

98.92

916.

0. 11

99.03

917.

0.11

99. 13

918.

0.11

99.24

919.

0.11

99.35

920.

0. 11

99.46

921.

0.11

99.57

922 •

0. 11

99.68

923.

0. 11

99.78

924.

0. 11

99.89

925.

0. 11

100.00

NUMBER OF SPECIES

15

NUMBER OF INDIVIDUALS

232.

149

CRUISE EX8001

STATION 34

GRAB 1

RANK SPECIES NAME

1 AGLAOPHAMUS NEOTENUS

2 SCOLOPLOS SP.

3 OUENIA FUSIFORMIS

4 NEOMYSIS AMERICANA

5 EUPORELLA TRUNCATULA

6 ARGISSA HAMATIPES

7 DIASTYLIS SCULPTA

8 COROPHIUM TUBERCULATUM

9 PHOTIS MACROCOXA

10 ERYTHROPS ERYTHROPHTHALMA

11 CEREBRATULUS LACTEUS

12 ARICIDEA SUECICA

13 AMPHARETE ARCTICA

14 YOLDIA LIMATULA

15 CYLICHNA ALBA

COUNT CUM COUNT

2

CUM 7.

172

172.

74. 14

74.14

20

192.

8.62

82.76

12

204.

5.17

87.93

11

215.

4.74

92.67

219.

1 .72

94.40

223.

1.72

96.12

224.

0.43

96.55

225.

0.43

96.98

226.

0.43

97.41

227.

0.43

97.84

228.

0.43

98.28

229.

0.43

98.71

230.

0.43

99.14

231.

0.43

99.57

232.

0.43

100.00

INDIVIDUALS PER M2

2320

CRUISE EX8001

STATION 35

RANK

SPECIES NAME

1

AGLAOPHAMUS NEOTENUS

2

OU/ENIA FUSIFORMIS

3

NEOMYSIS AMFRICANA

4

AMFELISCA AUDITA

5

EUHORELLA TRUNCATULA

6

MELITA N.SP.

7

CASCO PIGELOUI

8

MULINIA LATERALIS

9

NASSARIUS TRIVITTATUS

NUMKER OF SPECIES 9

NUMBER OF INDIVIDUALS 618.
INDIVIDUALS PER M2 6180

150

COUNT CUM COUNT

•/.

CUM 7.

519

519.

83.98

83.98

73

59?.

11 .81

95.79

16

608.

2.59

98.38

5

613.

0.81

99.19

614.

0.16

99.33

615.

0.16

99.51

616.

0.16

99.68

617.

0.16

99.84

618.

0.16

100.00

CRUISE EX8001

STATION 3A

RANK

1

SPECIES NAME
MEMERTEA P

AGLAOPHAMUS NEOTENUS
CORORHIUM INSIMOSUM
AMRELISCA AUDITA
CHIROTJOTEA COECA
PHOLOE MINUTA
NERHTYS BUCERA
PRIONOSriO STEENSTRUPI

NUMBER OF SPECIES 8

NUMBER OF INDIVIDUALS 12.
INMYIIiUALS PER M2 120

151

COUNT

CUM COUNT

7.

CUM '/.

3.

3.

25 . 00

25.00

3.

6.

25.00

50.00

7.

8.33

58.33

8.

0.33

46.67

9.

8.33

75.00

10.

8.33

83.33

11 .

8.33

91 .67

12.

8.33

100.00

CRUISE EX8001

STATION 37

RANK

SPECIES NAME

1

OLIGOCHAETA

2

BALANUS BALANOIDES

3

MYTILUS EPULIS

4

STREBLOSPIO BENEUICTI

5

POLYDORA LIGNI

6

THARYX SP.

7

JAERA SP.

8

LITTORINA LITTOREA

9

HETEROMASTUS FILIFORMIS

10

NEREIS VIRENS

11

GAMMARUS OCEANICUS

12

MACOMA BALTHICA

13

PYGOSPIO ELEGANS

14

MYA ARENARIA

15

PHOLOE M1NUTA

16

COROPHIUh TUBERCULATUM

17

CAPITELLA CAPITATA

18

COROPHIUM VOLUTATOR

19

ETEONE LONGA

20

POLYHORA SP.

21

NEMERTEA D

22

HYEIROBIA SP.

23

CALLIOSTOMA OCCIUENTALE

NUMBER OF SPECIES 23

NUMBER OF INBIVI HUALS 3638.
INDIVIDUALS PER M2 36380

152

COUNT

CUM COUNT

'/.

CUM •/.

1462.

1462.

40. 19

40.19

904.

2366.

24.85

65.04

508.

2874.

13.96

79.00

161.

3035.

4.43

83.42

150.

3185.

4.12

87.55

101 .

3286.

2.78

90.32

90.

3376.

2.47

92.80

87.

3463.

2.39

95.19

47.

3510.

1 .29

96.48

43.

3553.

1 . 18

97.66

15.

3568.

0.41

98.08

13.

3581 .

0.36

98.43

12.

3593.

0.33

98.76

8.

3601 .

0.22

98.98

7.

3608.

0.19

99. IB

5.

3613.

0. 14

99.31

5.

3618.

0. 14

99.45

4 .

3622.

0.11

99.56

4.

3626.

0.11

99.67

4.

3630.

0.11

9V.78

3.

3633.

0.08

99.86

3.

3636.

0.08

99.94

2.

3638.

0.05

100.00

CRUISE EX8001

STATION 38

RANK SPECIES NAME

1 AGLAOPHAMUS NEOTENUS

2 PRIONOSPIO STEENSTRUPI

3 MELITA N.SP.

4 OLIGOCHAETA

5 EUEiORELLA TRUNCATULA

6 CASCO BIGELOUI

7 METERYTHROPS ROBUSTA

8 ETEONE LONGA

9 MEDIOMASTUS AMBISETA

10 THARYX SP.

11 AMPHARETE ARCTICA

12 LACUNA VINCTA

NUMBER OF SPECIES

i:

NUMBER OF INDIVIDUALS 129.
INDIVIDUALS PER M2 1290

153

COUNl

CUM COUNT

•/.

CUM 7.

1 1 1

111 .

86.05

86.05

6

117.

4.65

90.70

2

119.

1 . 55

92.25

2

121.

1 .55

93.80

1 22 *

0.78

94.57

123.

0.78

95.35

124.

0.78

96.12

125.

0.78

96.90

126.

0.78

97.67

127.

0.78

98.45

128.

0.78

99.22

129.

0.78

100.00

CRUISE EXB001

STATION 3?

RANK SPECIES NAME

1 AGLAOPHAMUS NEOTENUS

2 PRIONOSPIO STEENSTRUPI

3 MELITA N.SP.

4 SCOLOPLOS SP.

5 MEDIOMASTUS AMBISETA

6 NINOE NIGRIPES

7 ARICIDEA JEFFREYSII

8 EUDORELLA TRUNCATULA

9 CEREBRATULUS LACTEUS

10 ARICIDEA SUECICA

11 ERYTHROPS ERYTHROPHTHALMA

12 NEOMYSIS AMERICANA

13 METERYTHROPS ROBUSTA

NUMBER OF SPECIES 13

NUMBER OF INDIVIDUALS 165.
INDIVIDUALS PER M2 1650

154

COUNT CUM COUNT

jj *
51 .
21.
19.

5.

4.

3.

2.

1.

1.

1 .

1.

1.

106.
127.
146.
151.
155.
158.
160.
161.
162.
163.
164.
165.

X

CUM

X

33

33

33

33

30

91

64

24

12

73

76

97

11

52

88

48

3

03

91

52

2

42

93

94

1

82

95

76

1

21

96

97

61

97

58

61

9ft

Ifl

61

98

7 9

61

99

39

61

100

00

CRUISE EX8001

STATION 40

NUMBER OF SPECIES

NUMBER 01 INDIVIDUALS 75.
INHIYItiUALS PER M2 750

155

RANK

1

2

3

4

5

6

7

8

9

10

1 1

12

13

14

15

SPECIES NAME
AGLAOPHAMUS NEOTENUS
ANEMONE A

ARICIHEA JEFFREYSII
PRIONOSPIO STEENSTRUPI
MELITA N.SP.
NINOE N I GRIPES
NEOMYSIS AMERICANA
YOLIHA LIMATULA
CEREBRATULUS LACTEUS
ARICIBEA SUECICA
ERYTHROPS ERYTHROPHTHALMA
ORCHOMENELLA PINGUIS
ARGISSA HAMATIPES
NUCULA PELPHINOUONTA
CERASTODERMA PINNULATUM

COUNT CUM COUNT

X

CUM V.

23

23.

30.67

30.67

15

38.

20.00

50.67

12

50.

16.00

66.67

9

59.

12.00

78.67

3

62.

4.00

82.67

•7

64.

2.67

85.33

2

66.

2.67

88.00

2

68.

2.67

90.67

69.

1 .33

92.00

70.

1.33

93.33

71 .

1 .33

94.67

72.

1 .33

96.00

73.

1 .33

97.33

74.

1 .33

98.67

75.

1.33

100.00

156

CRUISE EX8001

STATION 41

GRAB 1

RANK

1

2

3

4

5

6

7

8

9

10

11

12

13

14

15

16

17

18

19

20

21

22

23

24

25

26

27

28

29

SPECIES NAME
AMPELISCA ABUITA
NEPHTYS SP.
POLYTJORA LIGNI
PHOT IS MACROCOXA
MEIHOMASTUS AMBISETA
NASSARIUS TRIUITTATUS
TELLINA AGILIS
PRIONOSPIO STEENSTRUPI
DULICHIA MONOCANTHA
OLIGOCHAETA
ORCHOMENELLA PINGUIS
CIRRATULIBAE
EUBORELLA TRUNCATULA
OXYUROSTYLIS SMITHI
CAPITELLA CAPITATA
HETEROMASTUS EILIFORMIS
NINOE NIGRIF'ES
F'HOLOE MINUTA
PHERUSA AFFINIS
ANEMONE A
SCOLOF'LOS SP.
ARICIDEA JEFFREYSII
PHYLLOHOCE MACULATA
SYLLIHAE
ETEONE LONGA
MALHANOPSIS ELONGATA
MELITA N.SP.
MULINIA LATERALIS
CAMPANULARIA

COUN

CUM COUNT

y.

CUM V.

365

365.

45.17

45. 17

297

662.

36 . 76

81 .93

35

697.

4.33

86.26

21

718^

2.60

88.86

13

731.

1 .61

90.47

12

743.

1 .49

91 .96

10

753.

1 .24

93.19

6

759.

0. 74

93.94

6

765.

0.74

94.68

5

770.

0.62

95.30

5

775.

0.62

91.. 92

4

779.

0.50

96.41

4

783.

0.50

96.91

4

787.

0.50

97.40

3

790.

0.37

97.77

3

793.

0.37

98. 14

2

795.

0.25

98.39

?

797.

0.25

98.64

2

799.

0.25

98.89

800.

0.12

99.01

801 .

0.12

99. 13

802.

0. 12

99.26

803.

0.12

99.38

804.

0.12

99.50

805.

0.12

99.63

806.

0.1 7

99.75

807.

0.12

99.88

808.

0.12

100.00

NUMBER OF SPECIES

NUMBER OF INIUV I DUALS 808. +
INBIUIHUALS PER M2 8080+

CRUISE EX8001

STATION 12

RANK SPECIES NAME

1 AGLAOPHAMUS NEOTENUS

2 SPIRORBIS BOREALIS

3 DULICHIA MONOCANTHA

4 NASSARIUS TRIVITTATUS

5 NEPHTYS INC ISA

6 MULINIA LATERALIS

7 HYUROBIA SP.

8 [HASTYLIS SCULPTA

9 SPIRORBIS SP.

10 AEGININA LONGICORNIS

11 EUDORELLA TRUNCATULA

12 AMPELISCA ABDITA

13 ORCHOMENELLA PINGUIS

14 METERYTHROPS ROBUSTA

15 NEOMYSIS AMERICANA

16 ANEMONE A

17 POLYDORA LIGNI

18 NEREIS VIRENS

19 GEMMA GEMMA

NUMBER OF SPECIES

19

NUMBER OF INPIV I DUALS 272.
INDIVIDUALS PER M2 2720

157

COUNT CUM COUNT

■/.

CUM X

235

235.

86.40

86.40

6

241.

2.21

88.60

A

245.

1 .47

90.07

A

249.

1.47

91.54

3

252.

1. 10

92.65

3

255.

1 . 10

93.75

3

258.

1 .10

94.85

2

260.

0.74

95.59

2

262.

0.74

96.32

263.

0.37

96.69

264.

0.37

97.06

265.

0.37

97.43

266.

0.37

97.79

267.

0.37

98.16

268.

0.37

98.53

269.

0.37

98.90

270.

0.37

99.26

271.

0.37

99.63

272.

0.37

100.00

NUMBER OF SPECIES 7

NUMBER OF INDIVIDUALS 44.
INDIVIDUALS PER M2 440

158

CRUISE EX8001

STATION 43

GRAB 1

RANK.

SPECIES NAME

1

AGLAOPHAMUS NEOTENUS

5

NEOMYSIS AMERICANA

3

ANEMONE A

4

METERYTHROPS ROBUSTA

5

YOLDIA LIMATULA

6

MULINIA LATERALIS

7

NASSARIUS TRIVITTATUS

COUNT

CUM COUNT

X

CUM 7.

24.

24.

54.55

54 • 55

12.

36.

27.27

81.82

40.

9.09

90.91

41.

2.27

93.18

42.

2.27

95.45

43.

2 . 27

97.73

44.

2.27

100.00

159

CRUISE EX8001

STATION 44

CRAB 1

RANt, SPECIES NAME

1 AMPELISCA AUDITA

2 COROPHIUM CRASSICORNE

3 PRIONOSPIO STEENSTRUPI

4 LUMBRINERIS TENUIS

5 AGLAOPHAMUS NEOTENUS

6 ARICIDEA JEFFREYSII

7 PHOTIS MACROCOXA

8 MEDIOMASTUS AMBISETA

9 IiULICHIA MONOCANTHA

10 PHOXOCEPHALUS HOLBOLLI

11 ORCHOMENELLA PINGUIS

12 EUDORELLA TRUNCATULA

13 NINOE NIGRIPES

14 PHYLLOEIOCE MUCOSA

15 [HASTYLIS SCULPTA

16 PHOLOE MINUTA

17 ETEONE LONGA

18 AMPHARETE ACUTIFRONS

19 SCOLOPLOS SP.

20 NEPHTYS INC ISA

21 SACCOGLOSSUS KOUALEVSM I

22 POTAMILLA NEGLECTA

23 NEMERTEA A

24 OLIGOCHAETA

25 AMPHARETE ARCTICA

26 MACOMA BALTHICA

27 OUENIA FUSIFORMIS

28 LUMBRINERIS BREVIPES

29 ARGISSA HAMATIPES

30 ASABELLIDES OCULATA

31 STAURONEREIS CAECUS

32 MICROPHTHALMOS ABERRANS

33 POLYCIRRUS MEDUSA

34 PHYLLOHOCE MACULATA

35 METERYTHROPS ROBUSTA

36 DIASTYLIS POLITA

37 UNCI OLA IRRORATA

38 NEMERTEA H

39 CERIANTHUS BOREALIS

40 NUCULA DELPHINODONTA

41 TELLINA AGILIS

42 HYDROIO A

COUNT CUM COUNT

7.

CUM V.

563

563.

26.27

26.27

440

1003.

20.53

46.80

264

1267.

12.32

59.12

207

1474.

9.66

68.78

116

1590.

5.41

74.20

106

1696.

4.95

79.14

75

1771 .

3.50

82.64

51

1822.

2.38

85.02

46

1868.

2.15

87.17

45

1913.

2.10

89.27

38

1951 .

1 .77

91 .04

36

1987.

1.68

92.72

34

2021.

1 .59

94.31

27

2048.

1 .26

95.57

14

2062.

0.65

96.22

13

2075.

0.61

96.83

11

2086.

0.51

97.34

7

2093.

0.33

97.67

5

2098.

0.23

97.90

5

2103.

0.23

98.13

5

2108.

0.23

98.37

4

2112.

0.19

98.55

4

2116.

0. 19

98.74

3

2119.

0.14

98.88

3

21 22 .

0.14

99.02

3

2125.

0.14

99.16

2

2127.

0.09

99.25

2

2129.

0.09

99.35

2

2131 .

0.09

99.44

2132.

0.05

99.49

2133.

0.05

99.53

2134.

0.05

99.58

2135.

0.05

99.63

2136.

0.05

99.67

2137.

0.05

99.72

2138.

0.05

99.77

2139.

0.05

99.81

2140.

0.05

99.86

2141 .

0.05

99.91

2142.

0.05

99.95

2143.

0.05

100.00

NUMBER OF SPECIES 42

NUMBER OF INDIVIDUALS 2143. +
INDIVIDUALS PER M2 21430+

CRUISE EX8001

STATION 45

RANK
1

6
7
8
9
10
1 1
12
13
14
15
1*
17
18
19
20
21

SPECIES NAMF
AGLAOPHAMUS NEOTENUS
PRIONOSPIO STEENSTRUPI
ARICIBEA JEFFREYSII
THARYX SP.
NINOE NIGRIPES
NEREIS ylRENS
MEDIOMASTUS AMBISETA
COROPHIUM CRASSICORNE
SCOLOPLOS SR.
MICROPHTHALMOS ABERRANS
YOLDIA LIMATULA
CEREBRATULUS LACTEUS
OXYUROSTYLIS SMITHI
EUnORELLA TRUNCATULA
AMPELISCA ABDITA
F'HOXOCEF'HALUS HOLBOLLI
UNCI OLA IRRORATA
CASCO BIGELOUI
OLIGOCHAETA
ETEONE LONGA
NEFHTYS INCISA

NUMBER OF SRECIES

21

NUMBER OF INUI VIDUALS 675.
INDiyiDUALS PER M2 6750

160

COUNT CUM COUNT

•/.

CUM V.

406

406.

60. 15

60. 15

110

516.

16.30

76.44

55

571.

8.15

84.59

53

624.

7.85

92.44

13

637.

1 .93

94.37

9

646.

1 .33

95.70

6

652.

0.89

96.59

5

657.

0.74

97.33

4

661 .

0.59

97.93

3

664.

0.44

98.37

665.

0.15

98.52

666.

0. 15

98.67

667.

0. 15

98.81

668.

0.15

98.96

669.

0.15

99.11

670.

0. 15

99.26

671.

0. 15

99.41

672.

0.15

99.56

673.

0.15

99.70

674.

0.15

99.85

675.

0.15

100.00

CRUISE EX8001

STATION 46

GRAB 1

RANK SPECIES NAME

1 NUCULA DELPHINODONTA

2 LUMBBINERIS TENUIS

3 PHOXOCEPHALUS HOLBOLLI

4 EUDORELLA TRUNCATULA

5 PHOT IS MACROCOXA

6 PHYLLODOCE MUCOSA

7 PRIONOSPIO STEENSTRUPI

8 AMPELISCA ABDITA

9 NINOE NIGRIPES

10 ORCHOMENELLA PINGUIS

11 AGLAOPHAMUS NEOTENUS

12 DIASTYLIS SCULPTA

13 UNCIOLA IRRORATA

14 ARICIDEA JEFTREYSII

15 ETEONE LONGA

16 AMPHARETE ARCTICA

17 LEPTOCHEIRUS PINGUIS

18 OLIGOCHAETA

1? CEREBRATULUS LACTEUS

20 EDOTEA TRILOBA

21 NEMERTEA C

22 CERIANTHUS BOREALIS

23 COROPHIUM CRASSICORNE

24 CASCO BIGELOUI

25 MEDIOMASTUS AMBISETA

26 NEREIS VIRENS

27 PHYLLODOCE GROENLANDICA

28 SCOLOPLOS SP.

29 PHOLOE MINUTA

30 NEPHTYS INCISA

31 LUMBRINERIS FRAGILIS

32 GOLFINGIA VERRILLII

33 CANCER IRRORATUS

34 YOLIHA LIMATULA

35 MEMBRANIPORIHAE

36 MEMBRANIPORIDAE

COUNT CUM COUNT

X

CUM ■/.

503

503.

43.85

43.85

206

709.

17.96

61 .81

159

868.

13.86

75.68

42

910.

3.66

79.34

39

949.

3.40

82.74

28

977.

2.44

85.18

27

1004.

2.35

87.53

27

1031.

2.35

89.89

24

1055.

2.09

91.98

20

1075.

1 .74

93.72

11

1086.

0.96

94.68

11

1097.

0.96

95.64

6

1103.

0.52

96.16

5

1108.

0.44

96.60

4

1112.

0.35

96.95

4

1116.

0.35

97.30

4

1120.

0.35

97.65

3

1123.

0.26

97.91

3

1126.

0.26

98.17

3

1129.

0.26

98.43

2

1131.

0.17

98.60

2

1133.

0.17

98.78

2

1135.

0.17

98.95

2

1137.

0.17

99.13

1138.

0.09

99.22

1139.

0.09

99.30

1140.

0.09

99.39

1141 .

0.09

99.48

1142.

0.09

99.56

1143.

0.09

99.65

1144.

0.09

99.74

1145.

0.09

99.83

1146.

0.09

99.91

1147.

0.09

100.00

NUMBER OF SPECIES 36

NUMBER OF INDIVIDUALS 1147. +
INDIVIDUALS PER M2 11470 +

CRUISE EX8001

STATION 47

RANK SPECIES NAME

1 DIASTYLIS SCULF'TA

2 NASSARIUS TRIVITTATUS

3 LUMKRINERIS TENUIS

4 NINOE NIGRIPES

5 EUDORELLA TRUNCATULA

6 PHYLLODOCIDAE

7 AMPHIPHOLIS SGUAMATA

8 AMPHARETE ACUTIFRONS

9 FHERUSA AFFINIS

10 PAGURUS LONGICARPUS

11 AMPELISCA APDITA

12 LEF'TOCHEIRUS PINGUIS

13 UNCIOLA IRRORATA

14 ETEONE LONGA

15 NEREIS VIRENS

16 ARGISSA HAMATIPES

17 ANOMIA ACULEATA

18 CRENELLA DECUSSATA

19 ASTARTE UNDATA

20 CERASTOHERMA PINNULATUM

21 ASTERIAS SP.

22 MEMPRANIPORIDAE

23 MEMKRANIPORIDAE

NUMBER OF SPECIES

23

NUMBER OF INDIVIDUALS 80.+
INDIVIDUALS PER M2 800+

162

COUNT CUM COUNT

X

CUM y.

20

20.

25.00

25.00

1 1

31 .

13.75

38.75

9

40.

11.25

50.00

9

49.

11.25

61.25

5

54.

6.25

67.50

3

57.

3.75

71 .25

3

60.

3.75

75.00

T

62.

2.50

77.50

2

64.

2.50

80.00

T

66.

2.50

82.50

2

68.

2.50

85.00

-}

70.

2.50

87.50

o

72.

2.f.0

90.00

73.

1.25

91 .25

74.

1.25

92.50

75.

1.25

93.75

76.

1 .25

95.00

77.

1.25

96.25

78.

1 .25

97.50

79.

1 .25

98.75

80.

1.25

100.00

CRUISE EX8001

STATION 48

RANK SPECIES NAME

1 NUCULA UELPHINOtiONTA

2 PHOT IS MACROCOXA

3 EHOTEA TRILOBA

4 EUBORELLA TRUNCATULA

5 ORCHOMENELLA PINGUIS

6 IHASTYLIS SCULPTA

7 PHYLLOUOCE MUCOSA

8 AMPELISCA VADORUM

9 PHOXOCEPHALUS HOLBOLLI

10 LUMBRINERIS TENUIS

11 CRENELLA BECUSSATA

12 UNCIOLA IRRORATA

13 PRIONOSPIO STEENSTRUPI

14 COROPHIUM CRASSICORNE

15 HARPINIA PROPINQUA

16 ARICIEIEA JEFFREYSII

17 NINOE NIGRIPES

18 PITAR MORRHUANA

19 SCOLOPLOS SP.

20 LUMBRINERIS FRAGILIS

21 ANEMONE B

22 NUCULA ANNULATA

23 MYA ARENARIA

24 ECHINARACHNIUS PARMA

25 AMPHARETE ARCTICA

26 CERIANTHUS BOREALIS

27 PHILINE FINMARCHIA

28 CARBITA BOREALIS

29 NASSARIUS TRIVITTATUS

30 AGLAOPHAMUS NEOTENUS

31 PHOLOE MINUTA

32 PERIPLOMA PAPYRATIUM

33 MOIIIOLUS MOtHOLUS

34 LYONSIA HYALINA

35 LEPTOCHEIRUS PINGUIS

36 PHERUSA AFFINIS

37 SPIOPHANES BOMBYX

38 ETEONE LONGA

39 PHYLUM B

40 LINEUS RUBER

41 NEMERTEA E

42 CEREBRATULUS LACTEUS

43 NEMERTEA C

44 CERASTOBERMA PINNULATUM

45 MACOMA BALTHICA

46 ARGISSA HAMATIPES

NUMBER OF SPECIES 46

NUMBER OF INPIVIBUALS 1353.
INUIYICIUALS PER M2 13530

163

COUNl

CUM COUNT

X

cum y.

454.

454.

33.56

33.56

190

644.

14.04

47.60

91

735.

6.73

54.32

88

823.

6.50

60.83

86

909.

6.36

67.18

68

977.

5.03

72.21

61

1038.

4.51

76.72

44

1082.

3.25

79.97

39

1121.

2.88

82.85

36

1157.

2.66

85.51

23

1180.

1.70

87.21

23

1203.

1.70

88.91

21

1224.

1 .55

90.47

16

1240.

1.18

91 .65

15

1255.

1.11

92.76

13

1268.

0.96

93.72

13

1281.

0.96

94.68

7

1288.

0.52

95 . 20

6

1294.

0.44

95.64

6

1300.

0.44

96.08

4

1304.

0.30

96.38

4

1308.

0.30

96.67

4

1312.

0.30

96.97

3

1315.

0.22

97.19

3

1318.

0.22

97.41

3

1321.

0.22

97.63

3

1324.

0.22

97. B6

3

1327.

0.22

98.08

3

1330.

0.22

98.30

2

1332.

0.15

98.45

2

1334.

0.15

98.60

2

1336.

0.15

98.74

2

1338.

0.15

98.89

2

1340.

0.15

99.04

2

1342.

0.15

99.19

1343.

0.07

99.26

1344.

0.07

99.33

1345.

0.07

99.41

1346.

0.07

99.48

1347.

0.07

99.56

1348.

0.07

99.63

1349.

0.07

99.70

1350.

0.07

99.78

1351.

0.07

99.85

1352.

0.07

99.93

1353.

0.07

100.00

164

CRUISE EX8001

STATION 19

GRAB 1

RANK SPECIES NAME

1 PRIONOSPIO STEENSTRUPI

2 ECHINARACHNIUS PARMA

3 NUCULA DELPHINODONTA

4 UNCIOLA IRRORATA

5 POLYDORA QUADRILOBATA

6 NEMERTEA F

7 DODECACERIA SP.

8 PHYLLODOCE MACULATA

9 PHOLOE MINUTA

10 EDOTEA TRILOBA

11 NASSARIUS TRIVITTATUS

12 LUMBRINERIS TENUIS

13 PHOXOCEPHALUS HOLBOLLI

14 CRENELLA DECUSSATA

15 OLIGOCHAETA

16 AMPHIPHOLIS SOUAMATA

17 PHASCOLION STROMBI
IB ASTERIAS SP.

19 TEREIBELLIHAE

20 SYLLIS GRACILIS

21 POLYDORA SOCIALIS

22 AMPHARETIDAE

23 GATTYANA CIRROSA

24 HARMOTHOE EXTENUATA

25 OPHELINA ACUMINATA

26 LIMNORIA LIGNORUM

27 CANCER BOREALIS

28 PAGURUS ARCUATUS

29 MYA ARENARIA

30 CARDITA BOREALIS

31 OENOPOTA BICARINATA

32 SERTULARIA PUMILA

33 MEMBRANIPORA SP .

COUNT CUM COUNT

7.

CUM X

41

41 .

26. 11

26.11

17

58.

10.03

36.94

17

75.

10.83

47.77

11

86.

7.01

54.78

8

94.

5.10

59.87

6

100.

3.82

63.69

5

105.

3.18

66.88

5

110.

3. 18

70.06

5

115.

3.18

73.25

5

120.

3.18

76.43

5

125.

3.18

79.62

4

129.

2.55

82.17

4

133.

2.55

84.71

4

137.

2.55

87.26

3

140.

1.91

89. 17

2

142.

1.27

90.45

143.

0.64

91.08

144.

0.64

91.72

145.

0.64

92.36

146.

0.64

92.99

147.

0.64

93.63

148.

0.64

94.27

149.

0.64

94.90

150.

0.64

95.54

151.

0.64

96. 18

152.

0.64

96.82

153.

0.64

97.45

154.

0.64

98.09

155.

0.64

98.73

156.

0.64

99.36

157.

0.64

100.00

NUMBER OF SPECIES
NUMBER OF INDIVIDUALS
INDIVIDUALS PER M2

33
157. +

1570 +

CRUISE EX8001

STATION

RANK SPECIES NAME

1 POLYCIRRUS EXIMUS

2 AMPHARETE ARCTICA

3 NASSARIUS TRIVITTATUS

4 PHOXOCEPHALUS HOLBOLLI

5 ARCHIANNELIDA

6 PRIONOSPIO STEENSTRUPI

7 EUCLYMENE COLLARIS

8 OLIGOCHAETA

9 EXOGONE HEBES

10 CEREBRATULUS LACTEUS

11 POLYDORA SOCIALIS

12 NEMERTEA F

13 THARYX SP.

14 PHOLOE MINUTA

15 CHIRODOTEA COECA

16 SPHAEROSYLLIS ERINACEUS

17 SPIOPHANES BOMBYX

18 AGLAOPHAMUS CIRCINATA
1? HARMOTHOE EXTENUATA

20 OUENIIDAE

21 PHYLLODOCE MACULATA

22 PARAONIS LYRA

23 AMPHIPHOLIS SQUAMATA

24 ECHINARACHNIUS PARMA

25 PHILINE FINMARCHIA

26 AMPELISCA AGASSIZI

27 HALIMEDON SP.

28 ANONYX LILJEBORGI
2? UNCIOLA IRRORATA

30 EDOTEA TRILOBA

31 MEMBRANIPORIDAE

32 MEMBRANIPORIDAE

NUMBER OF SPECIES 32

NUMBER OF INDI VIDUALS 125. +
INDIVIDUALS PER M2 1250+

165

COUNT CUM COUNT

V.

CUM 7.

17

17.

13.60

13.60

12

29.

9.60

23.20

10

39.

8.00

31 .20

10

49.

8.00

39.20

9

58.

7.20

46.40

9

67.

7.20

53.60

7

74.

5.60

59.20

6

80.

4.80

64.00

6

86.

4.80

68.80

5

91.

4.00

72.80

5

96.

4.00

76.80

100.

3.20

80.00

104.

3.20

83.20

107.

2.40

85.60

110.

2.40

88.00

Ill .

0.80

88.80

112.

0.80

89.60

113.

0.80

90.40

114.

0.80

91.20

115.

0.80

92.00

116.

0.80

92.80

117.

0.80

93.60

118.

0.80

94.40

119.

0.80

95.20

120.

0.80

96.00

121 •

0.80

96.80

122.

0.80

97.60

123.

0.80

98.40

124.

0.80

99.20

125.

0.80

100.00

166

CRUISE EX8001

STATION

GRAB 1

RANK SPECIES NAME

1 LUMBRINERIS TENUIS

2 NUCULA DELPHINODONTA

3 NINOE NIGRIPES

4 PRIONOSPIO STEENSTRUPI

5 EUDORELLA TRUNCATULA

6 AGLAOPHAMUS NEOTENUS

7 CEREBRATULUS LACTEUS

8 ARICIDEA JEFFREYSII

9 DIASTYLIS SCULPTA

10 PITAR MORRHUANA

11 PHOXOCEPHAl.US HOLBOLLI

12 PARAONIS GRACILIS

13 AMPHARETE ARCTICA

14 MYA ARENARIA

15 PHYLLODOCE MUCOSA

16 NEPHTYS INCISA

17 AMPELISCA AGASSIZI

18 ARGISSA HAMATIPES

19 PHOLOE MINUTA

20 HARPINIA PROPINQUA

21 ASABELLIDES OCULATA

22 ETEONE LONGA

23 CERIANTHUS BOREALIS

24 PERIPLOMA PAPYRATIUM

25 ORCHOMENELLA PINGUIS

26 SCOLOPLOS SP .

27 ARCTICA ISLANPICA

28 PAGURUS LONGICARPUS

29 STENOPLEUSTES INERMIS

30 PHOTIS MACROCOXA

31 EUCLYMENE COLLARIS

32 ANEMONE A

33 NEMERTEA E

34 MODIOLUS MODIOLUS

35 YOLDIA LIMATULA

36 CERASTODERMA PINNULATUM

37 NASSARIUS TRIUITTATUS

COUNT

CUM COUNT

•/.

CUM 7.

145.

145.

27.46

27.46

92.

237.

17.42

44.89

B2.

319.

15.53

60.42

46.

365.

8.71

69.13

34.

399.

6.44

75.57

16.

415.

3.03

78.60

11.

426.

2.08

80.68

10.

436.

1.89

82.58

8.

444.

1.52

84.09

7.

451.

1.33

85.42

6.

457.

1.14

86.55

6.

463.

1 .14

87.69

6.

469.

1 .14

88.83

6.

475.

1.14

89.96

5.

480.

0.95

90.91

5.

485.

0.95

91 .86

4.

489.

0.76

92.61

4.

493.

0.76

93.37

4.

497.

0.76

94.13

3.

500.

0.57

94.70

3.

503.

0.57

95.26

3.

506.

0.57

95.83

3.

509.

0.57

96.40

3.

512.

0.57

96.97

2.

514.

0.38

97.35

2.

516.

0.38

97.73

2.

518.

0.38

98.11

1

519.

0.19

98.30

1 .

S20.

0.19

98.48

1

521.

0.19

98.67

1 .

522.

0.19

98.86

1 .

523.

0.19

99.05

1 ,

524.

0. 19

99.24

\ t

525.

0.19

99.43

1 ,

526.

0.19

99.62

I ,

527.

0.19

99.81

1 .

528.

0.19

100.00

NUMBER OF SPECIES 37

NUMBER OF INDIVIDUALS 528.

INDIVIDUALS PER M2

5280

CRUISE EX8001

STATION

RANK SPECIES NAME

1 PRIONOSPIO STEENSTRUPI

2 AGLAOPHAMUS NEOTENUS

3 ARICIDEA JEFFREYSII

4 LUMBRINERIS TENUIS

5 NUCULA UELPHINOIiONTA

6 PHYLLOBOCE MUCOSA

7 SCOLOPLOS SP.

8 EUBORELLA TRUNCATULA

9 NINOE NIGRIPES

10 OLIGOCHAETA

11 OUENIA FUSIFORMIS

12 PHOXOCEPHALUS HOLBOLLI

13 CEREBRATULUS LACTEUS

14 MEDIOMASTUS AMBISETA

15 AMPHARETE ARCTICA

16 ETEONE LONGA

17 NEPHTYS INC ISA

18 MODIOLUS MODIOLUS

19 PITAR MORRHUANA

20 NEMERTEA C

21 PHOLOE MINUTA

22 MALDANOPSIS ELONGATA

23 MACOMA BALTHICA

24 SOLEMYA BOREALIS

25 HYDROBIA SP .

26 AMPELISCA ABDITA

27 LEPTOCHEIRUS PINGUIS

28 NEOMYSIS AMERICANA

29 SERTULARIA PUMILA

NUMBER OF SPECIES 29

NUMHER OF INDIVIDUALS 761. +
INDIVIDUALS PER M2 7610+

167

COUNT

CUM COUNT

•/.

CUM ■/.

459

459.

60.32

60.32

79

538.

10.38

70.70

44

582.

5.78

76.48

35

617.

4.60

81 .08

32

649.

4.20

85.28

24

673.

3.15

88.44

23

696.

3.02

91 .46

19

715.

2.50

93.96

11

726.

1 .45

95.40

5

731 .

0.66

96.06

4

735.

0.53

96.58

1

738.

0.39

96.98

2

740.

0.26

97.24

9

742.

0.26

97.50

9

744.

0.26

97.77

2

746.

0.26

98.03

2

748.

0.26

98.29

o

750.

0.26

98.55

2

752.

0.26

98.82

753.

0.13

98.95

754.

0.13

99.08

i 75 j .

0.13

99.21

756.

0.13

99.34

757.

0.13

99.47

758.

0.13

99.61

759.

0. 13

99.74

760.

0.13

99.87

761.

0.13

100.00

168

CRUISE EX8001

STATION

GRAB 1

RANK SPECIES NAME

1 OLIGOCHAETA

2 PHOLOE MINUTA

3 ARICIHEA JEFFREYSII

4 PHOXOCEPHALUS HOLBOLLI

5 PRIONOSPIO STEENSTRUPI

6 MEDIOMASTUS AMBISETA

7 LUMBRINERIS TENUIS

8 THARYX SP.

9 SYLLIS CORNUTA

10 COROPHIUM INSIDIOSUM

11 ETEONE LONGA

12 ALBANIA ARENARIA

13 NUCULA DELPHINODONTA

14 AMPHIPHOLIS SQUAMATA

15 NINOE NIGRIPES

16 SCOLOPLOS ROBUSTUS

17 CRENELLA DECUSSATA

18 ASABELLIDES OCULATA

19 SCOLOPLOS SP.

20 NEREIS ZONATA

21 CEREBRATULUS LACTEUS

22 PHYLLOtiOCE MACULATA

23 SYLLIDAE

24 HARMOTHOE IMBRICATA

25 SYLLIS GRACILIS

26 CERIANTHUS BOREALIS

27 PHASCOLOPSIS GOULDII

28 COCCULINA SP.

29 PERIPLOMA PAPYRATIUM

30 MODIOLUS MODIOLUS

31 PANDORA GOULD I ANA

32 ASTARTE UNDATA

33 CARDITA BOREALIS

34 SKENEOPSIS PLANORBIS

35 CANCER IRRORATUS

36 HAPLOOPS TUB I COLA

37 LIMNORIA LIGNORUM

38 POLYCHAETE B

39 SPIRORBIS SP.

40 AUTOLYTUS SP.

41 EXOGONE HEBES
4 2 POLYDORA SP.

43 LEPIDONOTUS SQUAMATUS

44 NEPHTYS CILIATA

45 PHYLLODOCIDAE

4 6 MEMBRANIPORIDAE

47 CABEREA ELLISI

48 HYDROZOA

49 SERTULARIA PUMILA

COUNT

281

156

128

43

43

28

22

19

19

10

9

8

7

7

7

6

5

4

4

3

2

2

2

2

2

CUM COUNT
281 .
437.
565.
608.
651.
679.
701.
720.
739.
749.
758.
766.
773.
780.
787.
793.
798.
802.
806.
809.
811.
813.
815.
817.
819.
820.
821.
822.
823.
824.
825.
826.
827.
828.
829.
830.
831.
832.
833.
834.
835.
836.
837.
838.
839.

33.49

18
15

59
26
5. 13
5.13
3.34
2.62
2.26
2.26
1 .19
1.07
0.95
0.83
0.F13
0.83
0.72
0.60
0.48
0.48
0.36
0.24
0.24
0.24
0.24
0.24
0. 12
0.12
0.12
0.12
12
12
12
12
12
12
12

0.
0.
0.

0,

0.

0,

0.

0.12

0. 12

0.12

0.12

0.12

0.12

0.12

0.12

0. 12

cum y.

33.49
52.09
67.34
72.47
77.59
80.93
83.55
85.82
88.08
89.27
90.35
91.30
92.13
92.97
93.80
94.52
95.1 1
95.59
96.07
96.42
96.66
96.90
97. 14
97. 3B
97.62
97.74
97.85
97.97
98.09
98.21
98.33
98.45
98.57
98.69
98.81
98.93
99.05
99.17
99.28
99.40
99.52
99.64
99.76
99.88
100.00

NUMBER OF SPECIES
NUMBER OF INDIVIDUALS
INDIVIDUALS PER M2

49
839.+
8390 +

169

CRUISE EX8001

STATION

RANK
1

3
4

6

7

8

9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65

SPECIES NAME
PRI0N0SPI0 STEENSTRUPI
AMPELISCA AGASSIZI
NINOE N I GRIPES
SPIO FILICORNIS
MEDIOMASTUS AMBISETA
SCOLOPLOS SP.
LUMBRINERIS TENUIS
PHOT IS MACROCOXA
SABELLA PENICILLUS
THARYX SP.

NUCULA DELPHINODONTA
EUDORELLA TRUNCATULA
CASCO BIGELOUI
EDOTEA TRILOBA
ETEONE LONGA
ARGISSA HAMATIPES
DIASTYLIS SCULPTA
PHOXOCEPHALUS HOLBOLLI
CRENELLA DECUSSATA
CERIANTHUS BOREALIS
ASABELLIDES OCULATA
RHOEHNE LOVENI
PARAONIS GRACILIS
STENOPLEUSTES INERMIS
ORCHOMENELLA PINGUIS
LEPTOCHEIRUS PINGUIS
NEMERTEA K
NEMERTEA II
NEMERTEA I

CEREBRATULUS LACTEUS
AMPHARETE ACUTIFRONS
OLIGOCHAETA
PHYLLODOCE MUCOSA
STERNASPIS SCUTATA
PETALOSARSIA DECLIVIS
AMPELISCA MACROCEPHALA
CERASTODERMA PINNULATUM
PERIPLOMA PAPYRATIUM
NEMERTEA J

AGLAOPHAMUS NEOTENUS
DULICHIA MONOCANTHA
HIASTYLIS ABBREVIATA
MODIOLUS MODIOLUS
NUCULA ANNULATA
THYASIRA FLEXUOSA
PITAR MORRHUANA
NEMERTEA H
HARTMANIA MOOREI
HARMOTHOE IMBRICATA
AMPHARETE ARCTICA
PRAXILLELLA GRACILIS
SPIOPHANES BOMBYX
PHOLOE MINUTA
OPHELINA ACUMINATA
PHERUSA AFFINIS
GONIADA MACULATA
LAONICE CIRRATA
NEPHTYS INCISA
DIASTYLIS QUADRISPINOSA
CAMPYLASPIS RUBICUNDA
ANONYX LILJEBORGI
HAPLOOPS TUIUCDLA
MYTILUS EPULIS
ARCTICA ISLANHICA
YOLDIA LIMATULA

COUNT CUM COUNT

X

CUM 7.

1354

1354.

68.07

68.07

264

1618.

13.27

81.35

45

1663.

2.26

83.61

34

1697.

1 .71

85.32

28

1725.

1 .41

86.73

23

1748.

1 . 16

87.88

22

1770.

1.11

88.99

21

1791.

1 .06

90.05

16

1807.

0.80

90.85

12

1819.

0.60

91 .45

12

1831 .

0.60

92.06

11

1842.

0.55

92.61

11

1853.

0.55

93.16

9

1862.

0.45

93.61

7

1869.

0.35

93.97

6

1875.

0.30

94.27

5

1880.

0.25

94.52

5

1885.

0.25

94.77

5

1890.

0.25

95.02

1894.

0.20

95.22

1B98.

0.20

95.42

1902.

0.20

95.63

1906.

0.20

95.83

1910.

0.20

96.03

4

1914.

0.20

96.23

4

1918.

0.20

96.43

3

1921.

0.15

96.58

3

1924.

0.15

96.73

3

1927.

0. 15

96.88

3

1930.

0. 15

97.03

3

1933.

0.15

97.18

3

1936.

0.15

97.34

3

1939.

0. 15

97.49

3

1942.

0.15

97.64

3

1945.

0.15

97.79

3

1948.

0.15

97.94

3

1951.

0. 15

98.09

3

1954.

0.15

98.24

2

1956.

0.10

98.34

2

1958.

0.10

98.44

2

1960.

0.10

98.54

2

1962.

0.10

98.64

2

1964.

0. 10

98.74

2

1966.

0.10

98.84

2

1968.

0.10

98.94

2

1970.

0. 10

99.04

1971 .

0.05

99.09

1972.

0.05

99.14

1973.

0.05

99.20

1974.

0.05

99.25

1975.

0.05

99.30

1976.

0.05

99.35

1977.

0.05

99.40

1978.

0.05

99.45

1979.

0.05

99.50

1980.

0.05

99.55

1981.

0.05

99.60

1982.

0.05

99.65

1983.

0.05

99.70

1984.

0.05

99.75

1985.

0.05

99.80

1986.

0.05

99.85

1987.

0.05

99.90

1988.

0.05

99.95

1989.

0.05

100.00

NUMBER OF SPECIES

65

NUMBER OF INDIVIDUALS 1989.
INDIVIDUALS PER M2 19890

170

CRUISE EX8001

STATION 55

GRAB 1

RANK
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
IB
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53

SPECIES NAME
PRIONOSPIO STENNSTRUPI
MEDIOMASTUS AMBISETA
LEPTOCHEIRUS PINGUIS
LUMBRINERIS TENUIS
PHOXOCEPHALUS HOLBOLLI
EUDORELLA TRUNCATULA
HARPINIA PROPINQUA
ORCHOMENELLA PINGUIS
ETEONE LONGA
DIASTYLIS SCULPTA
ARICIDEA JEFFREYSII
CERIANTHUS BOREALIS
PHOT IS MACROCOXA
THARYX SP.
FHOLOE MINUTA
PHYLLODOCE MUCOSA
AGLAOPHAMUS NEOTENUS
NEREIS GRAYI
OLIGOCHAETA
POTAMILLA NEGLECTA
ARGISSA HAMATIPES
NINOE NIGRIPES
AMPELISCA AGASSIZI
PARAONIS GRACILIS
NUCULA DEUPHINODONTA
CEREBRATULUS LACTEUS
OUENIA FUSIFORMIS
CLYMENELLA TORGUATA
BRADA VILLOSA
OPHELINA ACUMINATA
COROPHIUM CRASSICORNE
ASABELLIDES OCULATA
NEREIDAE
PHERUSA AFFINIS
AHFHARETE ARCTICA
STAURONEREIS CAECUS
NEPHTYS INCISA
MODIOLUS MOBIOLUS
YOLDIA LIMATULA
NUCULA ANNULATA
CERASTODERMA FINNULATUM
NEMERTEA D
NEMERTEA C

LEPTOSTYLIS LONGIMANA
CASCO BIGELOWI
MELITA N.SP.
HARMOTHOE IMBRICATA
CAPITELLA CAFITATA
SCOLOPLOS SP.
LUMBRINERIS FRAGILIS
CARDITA BOREALIS
PERIPLOMA PAPYRATIUM
HYDROZOA

COUNT

CUM COUNT

2562.

2562.

119.

2681.

104.

2785.

71.

2856.

62.

2918.

37.

2955.

25.

2980.

24.

3004.

22.

3026.

20.

3046.

19.

3065.

16.

3081.

16.

3097.

14.

3111.

13.

3124.

13.

3137.

12.

3149.

11.

3160.

11 .

3171.

7.

3178.

6.

3184.

5.

3189.

4.

3193.

4.

3197.

4.

3201.

3.

3204.

3.

3207.

3.

3210.

3.

3213.

3.

3216.

2.

3218.

2.

3220.

2.

3222.

2.

3224.

2.

3226.

2.

3228.

2.

3230.

2.

3232.

2.

3234.

2.

3236.

2.

3238.

1.

3239.

1.

3240.

1.

3241.

1.

3242.

1.

3243.

1.

3244.

1.

3245.

1.

3246.

1.

3247.

1.

3248.

1.

3249.

+

X

78.86

3.66

3.20

2.19

91
14
77
74
68
62
0.58
0.49
0.49
0.43
0.40
0.40
0.37
0.34
0.34
0.22
0. 18
0.15
0.12
0.12
0.12
0.09
0.09
0.09
0.09
0.09
0.06
0.06
0.06
0.06
0.06
0.06
0.06
0.06
0.06
0.06
0.06
0.03
0.03
0.03
0.03
0.03
0.03
0.03
0.03
0.03
0.03
0.03

CUM X
78.86
82.52
85.72
87.90
89.81
90.95
91.72
92.46
93.14
93.75
94.34
94.83
95.32
95.75
96.15
96.55
96.92
97.26
97.60
97.81
98.00
98.15
98.28
98.40
98.52
98.61
98.71
98.80
98.89
98.98
99.05
99.11
99.17
99.23
99.29
99.35
99.41
99.48
99.54
99.60
99.66
99.69
99.72
99.75
99.78
99.82
99.85
99.88
99.91
99.94
99.97
100.00

NUMBER OF SPECIES

53

NUMBER OF INDIVIDUALS 3249.+

INDIVIDUALS PER M2

32490+

CRUISE EX8001

STATION 56

RANK SPECIES NAME

1 PRIONOSPIO STEENSTRUF'I

2 EUDORELLA TRUNCATULA

3 NINOE NIGRIPES

4 SCOLOFLOS SP.

5 NUCULA DELPHINODONTA

6 ERYTHROF'S ERYTHROPHTHALMA

7 NEPHTYS INCISA

8 CEREBRATULUS LACTEUS

9 DIASTYLIS SCULPTA

10 BATHYMEDON SP.

11 ARICIHEA SUECICA

12 MEDIOMASTUS AMBISETA

13 THARYX SP.

14 ARICIDEA JEFFREYSII

15 HARTMANIA MOORE I

16 LUMBRINERIS TENUIS

17 AGLAOPHAMUS NEOTENUS

18 YOLDIA LIMATULA

19 PITAR MORRHUANA

20 PHILINE FINMARCHIA

21 HAL I ME DON SP.

22 EUDORELLA HISPIDA

23 STENOPLEUSTES INERMIS

24 ARGISSA HAMATIPES

25 METOPELLA ANGUSTA

26 ORCHOMENELLA PINGUIS

27 METERYTHROPS ROBUSTA

NUMBER OF SPECIES 27

NUMBER OF INDIVIDUALS

INDIVIDUALS PER M2

2050

171

COUNT

CUM COUNT

7.

CUM X

92.

92.

44.88

44.88

41

133.

20.00

64.88

10

143.

4.88

69.76

8

151.

3.90

73.66

8

159.

3.90

77.56

8

167.

3.90

81.46

4

171.

1.95

83.41

4

175.

1.95

85.37

4

179.

1.95

87.32

4

183.

1.95

89.27

3

186.

1.46

90.73

2

188.

0.98

91.71

2

190.

0.98

92.68

2

192.

0.98

93.66

193.

0.49

94.15

194.

0.49

94.63

195.

0.49

95.12

196.

0.49

95.61

197.

0.49

96.10

198.

0.49

96.59

199.

0.49

97.07

200.

0.49

97.56

201.

0.49

98.05

202.

0.49

98.54

203.

0.49

99.02

204.

0.49

99.51

205.

0.49

100.00

CRUISE EX8001

STATION 57

RANK SPECIES NAME

1 PRIONOSPIO STEENSTRUPI

2 EUDORELLA TRUNCATULA

3 SCOLOPLOS SP.

4 NEPHTYS INCISA

5 CEREBRATULUS LACTEUS

6 ERYTHROPS ERYTHROPHTHALMA

7 MEDIOMASTUS AMBISETA

8 NUCULA DELPHINODONTA

9 METERYTHROPS ROBUSTA

10 ARGISSA HAMATIPES

11 PETALOPROCTUS TENUIS

12 ARICIDEA JEFFREYSII

13 NINOE NIGRIPES

14 NASSARIUS TRIVITTATUS

15 BATHYHEDON SP.

16 OUENIIHAE

17 YOLDIA LIMATULA

18 CERASTODERMA PINNULATUM
1? DIASTYLIS SCULPTA

20 AMPELISCA VADORUM

21 STENOPLEUSTES INERMIS

22 PARAONIS GRACILIS

23 LUMBRINERIS TENUIS

24 THARYX SP.

25 OLIGOCHAETA

NUMBER OF SPECIES 25

NUMBER OF INDIVIDUALS 236.
INDIVIDUALS PER M2 2360

172

COUNT CUM COUNT

X

CUM •/.

10?

109.

46. 19

46.19

53

162.

22.46

68.64

16

178.

6.78

7S.42

11

189.

4.66

80.08

6

195.

2.54

82.63

4

199.

1 .69

84 . 32

4

203.

1 .69

86.02

3

206.

1.27

87.29

3

209.

1 .27

88.56

3

212.

1.27

89.83

3

215.

1 .27

91. 10

3

218.

1.27

92.37

3

2">j

1 .27

93.64

2

223.

0.85

94.49

2

°25 «

0.85

95.34

2

227 .

0.85

96.19

228.

0.42

96.61

229.

0.42

97.03

230.

0.42

97.46

231.

0.42

97.88

232.

0.42

90.30

233.

0.42

98.73

234.

0.42

99.15

235.

0.42

99.58

236.

0.42

100.00

173

CRUISE EX8001

STATION 58

RANK SPECIES NAME

1 POLYPORA LIGNI

2 AGLAOPHAMUS NEOTENUS

3 AMPEL1SCA ABDITA

4 SCOLOPLOS SP.

5 POLY DORA SP.

6 TELLINA AGILIS

7 MEDIOMASTUS AMBISETA

8 ORCHOMENELLA PINGUIS

9 NEOMYSIS AMERICANA

10 OLIGOCHAETA

11 ASABELLIDES OCULATA

12 NASSARIUS TRIVITTATUS

13 STREBLOSPIO BENEDICTI

14 THARYX SP.

15 ETEONE LONGA

14 PHERUSA AFFINIS

17 MYTILUS EDULIS

18 MULINIA LATERALIS

19 GEMMA GEMMA

20 PARACAPRELLA TENUIS

21 DIASTYLIS SCULPTA

22 HALIMEDON SP.

23 DULICHIA MONOCANTHA

24 PHOXOCEPHALUS HOLBOLLI

COUNT CUM COUNT

V.

CUM ■/.

112

112.

24.14

24.14

109

221 •

23.49

47.63

77

298.

16.59

64.22

40

338.

8.62

72.84

39

377.

8.41

81.25

32

409.

6.90

88. 15

19

428.

4.09

92.24

9

437.

1.94

94.18

5

442.

1.08

95.26

4

446.

0.86

96.12

3

449.

0.65

96.77

3

452.

0.65

97.41

453.

0.22

97.63

454.

0.22

97.84

455.

0.22

98.06

456.

0.22

98.28

457.

0.22

98.49

458.

0.22

98.71

459.

0.22

98.92

460.

0.22

99.14

461.

0.22

99.35

462.

0.22

99.57

463.

0.22

99. 7B

464.

0.22

100.00

NUMBER OF SPECIES

24

NUMBER OF INDIVIDUALS 464.

INDIVIDUALS PER M2 4640

it U. S. GOVERNMENT PRINTING OFFICE: 1983--600-86 1--56".

(continued from inside front cover)

9. Phytoplankton Community Structure in Northeastern Coastal Waters of the
United States. II. November 1978. By Harold G. Marshall and Myra S. Cohn.
August 1981. Revised and reprinted October 1981. v + 14 p., 3 figs., 1 app.

10. Annual NEMP Report on the Health of the Northeast Coastal Waters of the
United States, 1980. Northeast Monitoring Program Report No. NEMP IV 81 A-H
0043. August 1981. Revised and reprinted January 1982. xxi + 79 p., 23
figs., 4 tables, 5 app.

11. Proceedings of the Summer Flounder (Paralichthys dentatus) Age and Growth
Workshop, 20-21 May 1980, Northeast Fisheries Center, Woods Hole,
Massachusetts. By Ronal W. Smith, Louise M. Dery, Paul G. Scarlett, and
Ambrose Jearld, Jr. December 1981. iv + 14 p., 10 figs., 6 tables.

12. Status of the Fishery Resources Off the Northeastern United States for

1981. By Resource Assessment Division, Northeast Fisheries Center. January

1982. iii + 114 p., 44 figs., 44 tables.

13. Gulf and Atlantic Survey for Selected Organic Pollutants in Finfish. By
Paul D. Boehm and Pam Hirtzer. April 1982. vii + 111 p., 46 figs., 31
tables, 2 app.

14. Ecosystem Definition and Community Structure of the Macrobenthos of the
NEMP Monitoring Station at Pigeon Hill in the Gulf of Maine. By Alan W.
Hulbert, Kenneth J. Pecci, Jonathan D. Witman, Larry G. Harris, James R.
Sears, and Richard A. Cooper. May 1982. xii + 143 p., 16 figs., 10 tables, 9
app.

15. Seasonal Phytoplankton Assemblages in Northeastern Coastal Waters of the
United States. By Harold G. Marshall and Myra S. Cohn. July 1982. vi + 31
p., 8 figs., 2 tables.

16. Contaminants in New York Bight and Long Island Sediments and Demersal
Species, and Contaminant Effects on Benthos, Summer 1980. By Robert N. Reid,
John E. O'Reilly, and Vincent S. Zdanowicz, eds. September 1982. x + 96 p.,
36 figs., 21 tables.

17. Summary of the Physical Oceanographic Processes and Features Pertinent to
Pollution Distribution in the Coastal and Offshore Waters of the Northeastern
United States, Virginia to Maine. By Merton C. Ingham (ed.), Reed S. Arm-
strong, J. Lockwood Chamberlin, Steven K. Cook, David G. Mountain, Ronald J.
Schlitz, James P. Thomas, James J. Bisagni, John F. Paul, and Catherine E.
Warsh. December 1982. vi + 166 p., 21 figs., 2 tables.

18. Stock Discrimination of Summer Flounder (Paralichthys dentatus) in the
Middle and South Atlantic Bights: Results of a Workshop. By Michael J.
Fogarty, Glenn DeLaney, John W. Gillikin, Jr., John C. Poole, Daniel E. Ralph,
Paul G. Scarlett, Ronal W. Smith, and Stuart J. Wilk. January 1983. iii +

14 p., 2 figs., 3 tables.

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